R. I. Amann, B. J. Binder, R. J. Olson, S. W. Chisholm, R. Devereux et al., 405 Combination of 16S rRNA-targeted oligonucleotide probes with flow cytometry for 406 analyzing mixed microbial populations, Appl. Environ. Microbiol, vol.56, pp.1919-1925, 1990.

J. P. Amend and E. L. Shock, Energetics of overall metabolic reactions of thermophilic and hyperthermophilic Archaea and Bacteria, FEMS Microbiology Reviews, vol.25, issue.2, pp.175-243, 2001.
DOI : 10.1111/j.1574-6976.2001.tb00576.x

I. Boutet, D. Jollivet, B. Shillito, D. Moraga, and A. Tanguy, Molecular identification of 410 differentially regulated genes in the hydrothermal-vent species Bathymodiolus thermophilus 411 and Paralvinella pandorae in response to temperature, BMC Genomics, vol.10, issue.222, p.412, 2009.

I. Boutet, R. Ripp, O. Lecompte, C. Dossat, E. Corre et al., 413 Conjugating effects of symbionts and environmental factors on gene expression in deep-sea 414 hydrothermal vent mussels, BMC Genomics, vol.12, issue.530, p.415, 2011.

R. Cannuel, P. G. Beninger, H. Mccombie, and P. Boudry, (Bivalvia: Mytilidae), The Biological Bulletin, vol.217, issue.2, pp.173-188, 2009.
DOI : 10.1086/BBLv217n2p173

C. M. Cavanaugh, Symbiotic chemoautotrophic bacteria in marine invertebrates from sulphide-rich habitats, Nature, vol.8, issue.5903, pp.58-61, 1983.
DOI : 10.1038/302058a0

C. M. Cavanaugh, S. L. Gardiner, M. L. Jones, H. W. Jannasch, and J. B. Waterbury, Prokaryotic Cells in the Hydrothermal Vent Tube Worm Riftia pachyptila Jones: Possible Chemoautotrophic Symbionts, prokaryotic 424 cells in the hydrothermal vent tube worm Riftia pachyptila Jones: possible 425 chemoautotrophic symbionts, pp.340-342, 1981.
DOI : 10.1126/science.213.4505.340

C. M. Cavanaugh, C. O. Wirsen, and H. W. Jannasch, Evidence for methylotrophic symbionts in a 427 hydrothermal vent mussel (bivalvia: mytilidae) from the mid-atlantic ridge, Appl. Environ. 428 Microbiol, vol.58, pp.3799-3803, 1992.

J. L. Charlou, J. P. Donval, Y. Fouquet, P. Jean-baptiste, and N. Holm, Geochemistry of high H2 and CH4 vent fluids issuing from ultramafic rocks at the Rainbow hydrothermal field (36??14???N, MAR), Chemical Geology, vol.191, issue.4, pp.345-359, 2002.
DOI : 10.1016/S0009-2541(02)00134-1

P. Chevaldonné, D. Desbruyères, and M. L. Haître, Time-series of temperature from three deep-sea hydrothermal vent sites, Deep Sea Research Part A. Oceanographic Research Papers, vol.38, issue.11, pp.1417-1430, 1991.
DOI : 10.1016/0198-0149(91)90014-7

J. J. Childress, C. R. Fisher, J. M. Brooks, M. C. Kennicutt, R. Bidigare et al., A 435 methanotrophic marine molluscan (bivalvia, mytilidae) symbiosis: mussels fueled by gas, Science, vol.233, pp.436-1306, 1986.

M. Segonzac, A review of the distribution of hydrothermal vent communities along 442 the northern Mid-Atlantic Ridge: dispersal vs. environmental controls, Hydrobiologia, vol.440, issue.443, pp.201-216, 2000.

A. Vangriesheim, Variations in deep-sea hydrothermal vent communities on the Mid- 447 Atlantic Ridge near the Azores plateau. Deep-Sea Res. I 48, pp.1325-1346, 2001.

D. L. Distel, H. K. Lee, and C. M. Cavanaugh, Intracellular coexistence of methano- and thioautotrophic bacteria in a hydrothermal vent mussel., Proceedings of the National Academy of Sciences, vol.92, issue.21, pp.9598-9602, 1995.
DOI : 10.1073/pnas.92.21.9598

N. Dubilier, R. Windoffer, and O. Giere, Ultrastructure and stable carbon isotope composition of the hydrothermal vent mussels Bathymodiolus brevior and B. sp. affinis brevior from the North Fiji Basin, western Pacific, Marine Ecology Progress Series, vol.165, pp.187-193, 1998.
DOI : 10.3354/meps165187

S. Duperron, The diversity of deep-sea mussels and their bacterial symbioses The Vent and Seep Biota, Topics in Geobiology, pp.137-167, 2010.

S. Duperron, C. Bergin, F. Zielinski, A. Blazejak, A. Pernthaler et al., A dual symbiosis shared by two mussel species, 458 Bathymodiolus azoricus and Bathymodiolus puteoserpentis (Bivalvia: Mytilidae), from 459 hydrothermal vents along the northern Mid-Atlantic Ridge, Environ. Microbiol, vol.457, issue.8, pp.1441-460, 2006.

S. Duperron, H. Guezi, S. M. Gaudron, P. Pop-ristova, F. Wenzhöfer et al., Relative abundances of methane- and sulphur-oxidising symbionts in the gills of a cold seep mussel and link to their potential energy sources, Geobiology, vol.382, issue.6, pp.481-491, 2011.
DOI : 10.1111/j.1472-4669.2011.00300.x

S. Duperron, M. C. Laurent, F. Gaill, and O. Gros, Sulphur-oxidizing extracellular bacteria in the gills of Mytilidae associated with wood falls, FEMS Microbiology Ecology, vol.63, issue.3, pp.338-349, 2008.
DOI : 10.1111/j.1574-6941.2008.00438.x

URL : https://hal.archives-ouvertes.fr/hal-00715244

S. Duperron, J. Lorion, S. Samadi, O. Gros, and F. Gaill, Symbioses between deep-sea mussels (Mytilidae: Bathymodiolinae) and chemosynthetic bacteria: diversity, function and evolution, Comptes Rendus Biologies, vol.332, issue.2-3, pp.298-310, 2009.
DOI : 10.1016/j.crvi.2008.08.003

URL : https://hal.archives-ouvertes.fr/hal-00750422

S. Duperron, T. Nadalig, J. C. Caprais, M. Sibuet, A. Fiala-medioni et al., Dual symbiosis in a Bathymodiolus sp. mussel from a methane seep on the Gabon 471 continental margin (southeast Atlantic): 16S rRNA phylogeny and distribution of the 472 symbionts in gills, Appl. Environ. Microbiol, vol.470, issue.71, pp.1694-1700, 2005.
URL : https://hal.archives-ouvertes.fr/hal-00152530

S. Duperron, M. Sibuet, B. J. Macgregor, M. M. Kuypers, C. R. Fisher et al., 474 Diversity, relative abundance and metabolic potential of bacterial endosymbionts in three 475 Bathymodiolus mussel species from cold seeps in the Gulf of Mexico, Environ. Microbiol, vol.476, issue.9, pp.1423-1438, 2007.

H. Felbeck, Chemoautotrophic Potential of the Hydrothermal Vent Tube Worm, Riftia pachyptila Jones (Vestimentifera), Science, vol.213, issue.4505, pp.336-338, 1981.
DOI : 10.1126/science.213.4505.336

J. J. Robinson and C. M. Cavanaugh, Ultrastructural, biochemical, and immunological 481 characterization of two populations of the mytilid mussel Bathymodiolus azoricus from the 482, 2002.

C. R. Fisher, J. M. Brooks, J. S. Vodenichar, J. M. Zande, J. J. Childress et al., The Co-occurrence of Methanotrophic and Chemoautotrophic Sulfur-Oxidizing Bacterial Symbionts in a Deep-sea Mussel, Marine Ecology, vol.23, issue.4, pp.277-289, 1993.
DOI : 10.1111/j.1439-0485.1993.tb00001.x

S. Halary, V. Riou, F. Gaill, T. Boudier, and S. Duperron, 3D FISH for the quantification of methane- and sulphur-oxidizing endosymbionts in bacteriocytes of the hydrothermal vent mussel Bathymodiolus azoricus, The ISME Journal, vol.34, issue.3, pp.284-292, 2008.
DOI : 10.2307/3593123

URL : https://hal.archives-ouvertes.fr/hal-00325687

K. S. Johnson, C. L. Beehler, C. M. Sakamoto-arnold, and J. J. Childress, In Situ Measurements of Chemical Distributions in a Deep-Sea Hydrothermal Vent Field, Science, vol.231, issue.4742, pp.1139-1141, 1986.
DOI : 10.1126/science.231.4742.1139

K. S. Johnson, J. J. Childress, C. L. Beehler, and C. M. Sakamoto, Biogeochemistry of 492 hydrothermal vent mussel communities: the deep-sea analogue to the intertidal zone, pp.993-1011, 1994.

H. D. Jones, O. G. Richards, and T. A. Southern, Gill dimensions, water pumping rate and body size in the mussel Mytilus edulis L., Journal of Experimental Marine Biology and Ecology, vol.155, issue.2, pp.213-237, 1992.
DOI : 10.1016/0022-0981(92)90064-H

E. Kádár, R. Bettencourt, V. Costa, R. S. Santos, A. Lobo-da-cunha et al., Experimentally induced endosymbiont loss and re-acquirement in the hydrothermal vent bivalve Bathymodiolus azoricus, Journal of Experimental Marine Biology and Ecology, vol.318, issue.1, pp.99-110, 2005.
DOI : 10.1016/j.jembe.2004.12.025

D. M. Karl, C. O. Wirsen, and H. W. Jannasch, Deep-Sea Primary Production at the Galapagos Hydrothermal Vents, Science, vol.207, issue.4437, pp.1345-1347, 1980.
DOI : 10.1126/science.207.4437.1345

L. Bris, N. Duperron, and S. , Chemosynthetic communities and biogeochemical energy 502 pathways along the Mid-Atlantic Ridge: the case of Bathymodiolus azoricus, p.503, 2010.

L. Bris, N. Zbinden, M. Gaill, and F. M. , Processes controlling the physico-chemical micro- 506 environments associated with Pompeii worms. Deep Sea Res. I 52 Coral Bleaching: Causes and Mechanisms Coral reefs: an ecosystem in transition, pp.1071-1083, 2005.

J. Lorion, S. Halary, J. Nascimento, . Do, S. Samadi et al., Evolutionary 510 history of Idas sp. Med (Bivalvia: Mytilidae), a cold seep mussel bearing multiple 511 symbionts, Cah. Biol. Mar, vol.53, pp.77-87, 2012.

J. M. Petersen, F. U. Zielinski, T. Pape, R. Seifert, C. Moraru et al., Hydrogen is an energy source for hydrothermal vent symbioses, Nature, vol.7, issue.7359, pp.2011-176
DOI : 10.1038/nature10325

URL : https://hal.archives-ouvertes.fr/cea-00903425

J. I. Prosser, Replicate or lie, Environmental Microbiology, vol.2, issue.7, pp.1806-1810, 2010.
DOI : 10.1111/j.1462-2920.2010.02201.x

A. M. Pruski and A. Fiala-médioni, Stimulatory effect of sulphide on thiotaurine synthesis in three hydrothermal-vent species from the East Pacific Rise, 518 R Development Core Team, 2013. R Development Core Team (2013). R: A language and 519 environment for statistical computing. R Foundation for Statistical Computing, pp.2923-2930, 2003.
DOI : 10.1242/jeb.00513

URL : https://hal.archives-ouvertes.fr/hal-00131173

V. Riou, S. Halary, S. Duperron, S. Bouillon, M. Elskens et al., Influence of CH<sub>4</sub> and H<sub>2</sub>S availability on symbiont distribution, carbon assimilation and transfer in the dual symbiotic vent mussel <i>Bathymodiolus azoricus</i>, Biogeosciences, vol.5, issue.6, pp.1681-1691, 0528.
DOI : 10.5194/bg-5-1681-2008

B. Shillito, F. Gaill, and J. Ravaux, The Ipocamp pressure incubator for deep-sea fauna, 0530.

B. Shillito, G. Hamel, C. Duchi, D. Cottin, J. Sarrazin et al., Live capture of megafauna from 2300 m depth, using a newly designed pressurized 533 recovery device, Deep Sea Res. I, vol.532, issue.55, pp.881-889, 2008.

E. Dinsdale, L. Kelly, and F. Rohwer, Metagenomic analysis of stressed coral 536 holobionts, Environ. Microbiol, vol.11, pp.2148-2163, 2009.

J. L. Trask and C. L. Van-dover, sp.) from the Lucky Strike hydrothermal vent field, Mid- Atlantic Ridge, Limnology and Oceanography, vol.44, issue.2, pp.334-343, 0539.
DOI : 10.4319/lo.1999.44.2.0334

C. Van-dover, The ecology of deep-sea hydrothermal vents, 2000.

C. L. Van-dover, C. R. German, K. G. Speer, L. M. Parson, and R. C. Vrijenhoek, Evolution and Biogeography of Deep-Sea Vent and Seep Invertebrates, Science, vol.295, issue.5558, pp.1253-1257, 2002.
DOI : 10.1126/science.1067361

V. Cosel, R. Cosel, R. V. Comtet, T. Krylova, and E. M. , Bathymodiolus (Bivalvia: Mytilidae) 545 from hydrothermal vents on the Azores Triple Junction and the Logatchev Hydrothermal 546 Field, Mid-Atlantic Ridge, Veliger, vol.42, pp.218-248, 1999.

W. Vesicomya, Isorropodon and Callogonia in the eastern Atlantic and the 549 Mediterranean, Sarsia, vol.86, pp.333-366

G. Wallner, R. Amann, and W. Beisker, Optimizing fluorescent in situ hybridization with rRNA- 551 targeted oligonucleotide probes for flow cytometric identification of microorganisms, Cytometry, vol.552, issue.14, pp.136-143, 1993.

C. Wentrup, A. Wendeberg, M. Schimak, C. Borowski, and N. Dubilier, Forever competent: deep-sea bivalves are colonized by their chemosynthetic symbionts throughout their lifetime, Environmental Microbiology, vol.12, issue.12, pp.3699-3713, 2014.
DOI : 10.1111/1462-2920.12597