J. Bascompte, P. Jordano, C. J. Meli, and J. M. Olesen, The nested assembly of plant-animal mutualistic networks, Proc. Natl Acad. Sci. USA, pp.9383-9387, 2003.
DOI : 10.1073/pnas.1633576100

J. Bascompte, P. Jordano, and J. M. Olesen, Asymmetric Coevolutionary Networks Facilitate Biodiversity Maintenance, Science, vol.312, issue.5772, pp.431-433, 2006.
DOI : 10.1126/science.1123412

U. Bastolla, M. A. Fortuna, A. Pascual-garc-ia, A. Ferrera, B. Luque et al., The architecture of mutualistic networks minimizes competition and increases biodiversity, Nature, vol.96, issue.7241, pp.1018-1020, 2009.
DOI : 10.1038/nature07950

C. Bernhardsson and P. K. Ingvarsson, L., Salicaceae), Molecular Ecology, vol.65, issue.9, pp.2197-2207, 2012.
DOI : 10.1111/j.1365-294X.2012.05524.x

C. Bernhardsson, K. M. Robinson, I. N. Abreu, S. Jansson, B. R. Albrectsen et al., Geographic structure in metabolome and herbivore community co-occurs with genetic structure in plant defence genes, Ecology Letters, vol.338, issue.6, pp.791-798, 2013.
DOI : 10.1111/ele.12114

D. A. Bohan, A. Raybould, C. Mulder, G. Woodward, A. Tamaddoni-nezad et al., Networking Agroecology, Adv. Ecol. Res, vol.49, pp.1-67, 2013.
DOI : 10.1016/B978-0-12-420002-9.00001-9

E. Canard, N. Mouquet, L. Marescot, K. J. Gaston, D. Gravel et al., Emergence of Structural Patterns in Neutral Trophic Networks, PLoS ONE, vol.7, issue.8, p.38295, 2012.
DOI : 10.1371/journal.pone.0038295.s005

M. F. Cattin, L. F. Bersier, C. Banasek-richter, R. Baltensperger, and J. P. Gabriel, Phylogenetic constraints and adaptation explain food-web structure, Nature, vol.427, issue.6977, pp.835-839, 2004.
DOI : 10.1038/nature02327

I. Chong and C. H. , Performance of some variable selection methods when multicollinearity is present, Chemometrics and Intelligent Laboratory Systems, vol.78, issue.1-2, pp.103-112, 2005.
DOI : 10.1016/j.chemolab.2004.12.011

J. E. Cohen, F. Briand, and C. M. Newman, Community food webs: data and theory, 1990.
DOI : 10.1007/978-3-642-83784-5

B. Dalsgaard, K. Trojelsgaard, A. M. Gonz-alez, D. Nogu-es-bravo, J. Ollerton et al., Historical climate-change influences modularity and nestedness of pollination networks, Ecography, vol.143, issue.12, pp.1331-1340, 2013.
DOI : 10.1111/j.1600-0587.2013.00201.x

C. F. Dormann and R. Strauss, A method for detecting modules in quantitative bipartite networks, Methods in Ecology and Evolution, vol.103, issue.3218, pp.90-98, 2013.
DOI : 10.1111/2041-210X.12139

C. F. Dormann, B. Gruber, and J. Fruend, Introducing the bipartite Package: analysing Ecological Networks, R. News, vol.8, pp.8-11, 2008.

C. F. Dormann, J. Fruend, N. Bluethgen, and B. Gruber, Indices, Graphs and Null Models: Analyzing Bipartite Ecological Networks, The Open Ecology Journal, vol.2, issue.1, pp.7-24, 2009.
DOI : 10.2174/1874213000902010007
URL : http://citeseerx.ist.psu.edu/viewdoc/summary?doi=10.1.1.323.9930

L. Eriksson, J. L. Hermens, E. Johansson, H. J. Verhaar, and S. Wold, Multivariate analysis of aquatic toxicity data with PLS, Aquatic Sciences, vol.20, issue.138, pp.217-241, 1995.
DOI : 10.1007/BF00877428

P. P. Feeny, Plant apparency and chemical defense Biochemical interaction between plants and insects, pp.1-40, 1976.

C. Fontaine, P. R. Guimar~-aes, S. Efi, N. Loeuille, J. Memmott et al., The ecological and evolutionary implications of merging different types of networks, Ecology Letters, vol.20, issue.11, pp.1170-1181, 2011.
DOI : 10.1111/j.1461-0248.2011.01688.x
URL : https://hal.archives-ouvertes.fr/bioemco-00644662

P. R. Guimar~-aes, V. Rico-gray, P. S. Oliveira, T. J. Izzo, S. F. Reis et al., Interaction Intimacy Affects Structure and Coevolutionary Dynamics in Mutualistic Networks, Current Biology, vol.17, issue.20, pp.1797-1803, 2007.
DOI : 10.1016/j.cub.2007.09.059

T. C. Ings, J. M. Montoya, J. Bascompte, N. Bl?-uthgen, L. Brown et al., Review: Ecological networks - beyond food webs, Journal of Animal Ecology, vol.3, issue.1, pp.253-269, 2009.
DOI : 10.1111/j.1365-2656.2008.01460.x

P. Jordano, J. Bascompte, and J. M. Olesen, Invariant properties in coevolutionary networks of plant-animal interactions, Ecology Letters, vol.99, issue.1, pp.69-81, 2003.
DOI : 10.1046/j.1461-0248.2003.00403.x

A. E. Krause, K. A. Frank, D. M. Mason, R. E. Ulanowicz, and W. W. Taylor, Compartments revealed in food-web structure, Nature, vol.426, issue.6964, pp.282-285, 2003.
DOI : 10.1038/nature02115

A. Krishna, P. R. Guimar~-aes-jr, P. Jordano, and J. Bascompte, A neutral-niche theory of nestedness in mutualistic networks, Oikos, vol.116, issue.11, pp.1609-1618, 2008.
DOI : 10.1111/j.1600-0706.2008.16540.x

T. Landelius, W. Josefsson, and T. Persson, A system for modelling solar radiation parameters with mesoscale spatial resolution, 2001.

S. Lavergne, N. Mouquet, W. Thuillier, and O. Ronce, Biodiversity and Climate Change: Integrating Evolutionary and Ecological Responses of Species and Communities, Annual Review of Ecology, Evolution, and Systematics, vol.41, issue.1, pp.321-350, 2010.
DOI : 10.1146/annurev-ecolsys-102209-144628

T. Lewinsohn and P. I. Prado, Structure in plant-animal interaction assemblages, Oikos, vol.113, issue.1, pp.174-184, 2006.
DOI : 10.1111/j.0030-1299.2006.14583.x

N. Loeuille and M. A. Leibold, Ecological consequences of evolution in plant defenses in a metacommunity, Theoretical Population Biology, vol.74, issue.1, pp.34-45, 2008.
DOI : 10.1016/j.tpb.2008.04.004

N. Loeuille and M. Loreau, Evolutionary emergence of size-structured food webs, Proc. Natl Acad. Sci. USA, pp.5761-5766, 2005.
DOI : 10.1073/pnas.0408424102

N. Loeuille and M. Loreau, Emergence of complex food web structure in community evolution models, Community Ecology, pp.163-179, 2009.
DOI : 10.1093/acprof:oso/9780199228973.003.00013

N. Loeuille, S. Barot, E. Georgelin, G. Kylafis, and C. Lavigne, Eco-Evolutionary Dynamics of Agricultural Networks, Adv. Ecol. Res, vol.49, pp.339-435, 2013.
DOI : 10.1016/B978-0-12-420002-9.00006-8

V. Luquez, D. Hall, B. R. Albrectsen, J. Karlsson, P. Ingvarsson et al., Natural phenological variation in aspen (Populus tremula): the SwAsp collection, Tree Genetics & Genomes, vol.16, issue.2, pp.279-292, 2008.
DOI : 10.1007/s11295-007-0108-y

N. Marron, S. Y. Dillen, and R. Ceulemans, Evaluation of leaf traits for indirect selection of high yielding poplar hybrids, Environmental and Experimental Botany, vol.61, issue.2, pp.103-116, 2007.
DOI : 10.1016/j.envexpbot.2007.04.002

C. F. Mcquaid and N. F. Britton, Host???parasite nestedness: A result of co-evolving trait-values, Ecological Complexity, vol.13, pp.53-59, 2013.
DOI : 10.1016/j.ecocom.2013.01.001

J. M. Montoya and R. V. Sol-e, Small World Patterns in Food Webs, Journal of Theoretical Biology, vol.214, issue.3, pp.405-412, 2002.
DOI : 10.1006/jtbi.2001.2460

J. M. Montoya, S. L. Pimm, and R. V. Sol-e, Ecological networks and their fragility, Nature, vol.27, issue.7100, pp.259-264, 2006.
DOI : 10.1038/nature04927

H. M?-uller-sch?-arer, U. Schaffner, and T. Steinger, Evolution in invasive plants: implications for biological control, Trends in Ecology & Evolution, vol.19, issue.8, pp.417-422, 2004.
DOI : 10.1016/j.tree.2004.05.010

J. Norberg, M. C. Urban, M. Vellend, C. A. Klausmeier, and N. Loeuille, Eco-evolutionary responses of biodiversity to climate change, Nature Climate Change, vol.12, issue.10, pp.747-751, 2012.
DOI : 10.1371/journal.pone.0000193

S. L. Nuismer, P. Jordano, and J. Bascompte, COEVOLUTION AND THE ARCHITECTURE OF MUTUALISTIC NETWORKS, Evolution, vol.8, issue.1, pp.338-354, 2012.
DOI : 10.1111/j.1558-5646.2012.01801.x

J. Ollerton, Biological Barter " : patterns of specialization compared across different mutualisms, pp.411-435, 2005.

M. Waser and J. Ollerton, Plant-pollinator interactions, from specialization to generalization

J. Ollerton, S. D. Johnson, L. Cranmer, and S. Kellie, The Pollination Ecology of an Assemblage of Grassland Asclepiads in South Africa, Annals of Botany, vol.92, issue.6, pp.807-834, 2003.
DOI : 10.1093/aob/mcg206

T. Poisot, E. Canard, N. Mouquet, and M. E. Hochberg, A comparative study of ecological specialization estimators, Methods in Ecology and Evolution, vol.11, issue.3, pp.537-544, 2012.
DOI : 10.1111/j.2041-210X.2011.00174.x

R. Team, R: a language and environment for statistical computing. R Foundation for Statistical Computing, 2013.

. Albrectsen, Genetic variation in functional traits influences arthropod community composition in aspen (Populus tremula L.) D, PLoS ONE, vol.7, p.37679, 2012.

K. M. Robinson, C. Hauzy, N. Loeuille, and B. R. Albrectsen, SwAspNet package: data from " Relative impacts of environmental variation and evolutionary history on the nestedness and modularity of tree-herbivore networks, Ecol. Evol, 2015.

T. N. Romanuk, Y. Zhou, U. Brose, E. L. Berlow, R. J. Williams et al., Predicting invasion success in complex ecological networks, Philosophical Transactions of the Royal Society B: Biological Sciences, vol.87, issue.20, pp.1743-1754, 2009.
DOI : 10.1073/pnas.192448799

G. Sanchez, E. , and C. Fontaine, Package 'DiscriMiner'. http://cran. rproject.org Stability of ecological communities and the architecture of mutualistic and trophic networks, Science, vol.329, pp.853-856, 2010.

A. R. Thompson, T. C. Adam, K. M. Hultgren, and C. E. Thacker, Ecology and Evolution Affect Network Structure in an Intimate Marine Mutualism, The American Naturalist, vol.182, issue.2, pp.58-72, 2013.
DOI : 10.1086/670803

F. S. Valdovinos, P. Moisset-de-espan-es, J. D. Flores, and R. Ramos-jiliberto, Adaptive foraging allows the maintenance of biodiversity of pollination networks, Oikos, vol.38, issue.6, pp.907-917, 2012.
DOI : 10.1111/j.1600-0706.2012.20830.x

F. J. Van-veen, C. B. Mueller, J. K. Pell, and H. C. Godfray, Food web structure of three guilds of natural enemies: predators, parasitoids and pathogens of aphids, Journal of Animal Ecology, vol.87, issue.1, pp.191-200, 2008.
DOI : 10.1111/j.1365-2656.2007.01325.x

D. P. Vazquez, C. J. Melian, N. M. Williams, N. Bl?-uthgen, B. R. Krasnov et al., Species abundance and asymmetric interaction strength in ecological networks, Oikos, vol.67, issue.7, pp.1120-1127, 2007.
DOI : 10.1111/j.0030-1299.2007.15828.x

V. Viswanath, B. R. Albrectsen, and S. H. Strauss, Global regulatory burden for field testing of genetically modified trees, Tree Genetics & Genomes, vol.5, issue.2, pp.221-226, 2012.
DOI : 10.1007/s11295-011-0445-8

T. G. Whitham, W. P. Young, G. D. Martinsen, C. A. Gehring, J. A. Schweitzer et al., COMMUNITY AND ECOSYSTEM GENETICS: A CONSEQUENCE OF THE EXTENDED PHENOTYPE, Ecology, vol.84, issue.3, pp.559-573, 2003.
DOI : 10.2307/2409439

R. J. Williams and N. D. Martinez, Simple rules yield complex food webs, Nature, vol.81, issue.6774, pp.180-183, 2000.
DOI : 10.1038/35004572

S. Wold, C. Albano, W. J. Dunn, I. , K. Esbensen et al., Multivariate data analysis: converting chemical data tables to plots, Intell. Instrum. Comput, vol.9, pp.197-216, 1989.

R. Wu and R. F. Stettler, Quantitative genetics of growth and development in Populus. III. Phenotypic plasticity of crown structure and function, Heredity, vol.88, issue.3, pp.299-310, 1998.
DOI : 10.1038/sj.hdy.6880860