C. S. Elton, The ecology of invasions by animals and plants, 1958.

J. L. Molnar, R. L. Gamboa, C. Revenga, and M. D. Spalding, Assessing the global threat of invasive species to marine biodiversity, Frontiers in Ecology and the Environment, vol.37, issue.9, pp.485-492, 1890.
DOI : 10.1023/B:BINV.0000034598.28718.2e

J. T. Carlton, Pattern, process, and prediction in marine invasion ecology, Biological Conservation, vol.78, issue.1-2, pp.97-106, 1996.
DOI : 10.1016/0006-3207(96)00020-1

D. Haydar, G. Hoarau, J. L. Olsen, W. T. Stam, and W. J. Wolff, Introduced or glacial relict? Phylogeography of the cryptogenic tunicate Molgula manhattensis (Ascidiacea, Pleurogona), Diversity and Distributions, vol.20, issue.1, pp.68-80, 2011.
DOI : 10.1111/j.1472-4642.2010.00718.x

J. P. Wares, D. S. Goldwater, B. Y. Kong, and C. W. Cunningham, Refuting a controversial case of a human-mediated marine species introduction, Ecology Letters, vol.55, issue.4, pp.577-584, 2002.
DOI : 10.1007/s00227-002-0780-2

J. W. Chapman, J. T. Carlton, M. R. Bellinger, and A. M. Blakeslee, Premature refutation of a human-mediated marine species introduction: the case history of the marine snail Littorina littorea in the northwestern Atlantic, Biological Invasions, vol.1, issue.6, pp.737-750, 2007.
DOI : 10.1007/s10530-006-9073-x

D. Haydar, What is natural? The scale of cryptogenesis in the North Atlantic Ocean, Diversity and Distributions, vol.19, issue.2, 2012.
DOI : 10.1111/j.1472-4642.2011.00863.x

N. Shenkar and B. J. Swalla, Global Diversity of Ascidiacea, PLoS ONE, vol.261, issue.6889, 2011.
DOI : 10.1371/journal.pone.0020657.s001

Z. Hoshino and T. Nishikawa, Taxonomic Studies of Ciona intestinalis (L.) and its allies, Publ. Seto Mar. Biol. Lab, vol.30, pp.61-79, 1985.

M. M. Suzuki, T. Nishikawa, and A. Bird, Genomic Approaches Reveal Unexpected Genetic Divergence Within Ciona intestinalis, Journal of Molecular Evolution, vol.84, issue.5, pp.627-635, 2005.
DOI : 10.1007/s00239-005-0009-3

L. Caputi, Cryptic speciation in a model invertebrate chordate, Proc. Natl. Acad. Sci. of USA, pp.9364-93690610158104, 2007.
DOI : 10.1073/pnas.0610158104

M. L. Nydam and R. G. Harrison, Genealogical relationships within and among shallow-water Ciona species (Ascidiacea), Marine Biology, vol.58, issue.5, pp.1839-1847, 2007.
DOI : 10.1007/s00227-007-0617-0

Z. Hoshino and T. Tokioka, An unusually robust Ciona from the Northeastern coast of Honsyu Island, Japan. Publ. Seto Mar. Biol. Lab, vol.15, pp.275-290, 1967.

R. H. Millar and L. M. In, Memoirs of typical British marine palnts and animals, XXXV, J. S. Colman), vol.123, 1953.

R. Pennati, Morphological Differences between Larvae of the Ciona intestinalis Species Complex: Hints for a Valid Taxonomic Definition of Distinct Species, PLOS ONE, vol.15, issue.5, 2015.
DOI : 10.1371/journal.pone.0122879.s005

A. Sato, N. Satoh, and J. D. Bishop, Field identification of ???types??? A and B of the ascidian Ciona intestinalis in a region of sympatry, Marine Biology, vol.19, issue.7, pp.1611-1619, 2012.
DOI : 10.1007/s00227-012-1898-5

M. L. Nydam and R. G. Harrison, Polymorphism and divergence within the ascidian genus Ciona, Molecular Phylogenetics and Evolution, vol.56, issue.2, pp.718-726042, 2010.
DOI : 10.1016/j.ympev.2010.03.042

C. Roux, G. Tsagkogeorga, N. Bierne, and N. Galtier, Crossing the Species Barrier: Genomic Hotspots of Introgression between Two Highly Divergent Ciona intestinalis Species, Molecular Biology and Evolution, vol.30, issue.7, pp.1574-1587, 2013.
DOI : 10.1093/molbev/mst066

A. Zhan, H. J. Macisaac, and M. E. Cristescu, Invasion genetics of the Ciona intestinalis species complex: from regional endemism to global homogeneity, Molecular Ecology, vol.11, issue.21, pp.4678-4694, 2010.
DOI : 10.1111/j.1365-294X.2010.04837.x

S. Bouchemousse, L. Lévêque, G. Dubois, and F. Viard, Co-occurrence and reproductive synchrony do not ensure hybridization between an alien tunicate and its interfertile native congener, Evolutionary Ecology, vol.19, issue.433, pp.69-87, 2016.
DOI : 10.1007/s10682-015-9788-1
URL : https://hal.archives-ouvertes.fr/hal-01227971

C. C. Lambert and G. Lambert, Non-indigenous ascidians in southern California harbors and marinas, Marine Biology, vol.130, issue.4, pp.675-688, 1998.
DOI : 10.1007/s002270050289

T. W. Therriault and L. Herborg, Predicting the potential distribution of the vase tunicate Ciona intestinalis in Canadian waters: informing a risk assessment, ICES Journal of Marine Science, vol.65, issue.5, pp.788-794, 2008.
DOI : 10.1093/icesjms/fsn054

A. Zhan, Complex genetic patterns in closely related colonizing invasive species, Ecology and Evolution, vol.19, issue.7, pp.1331-1346, 2012.
DOI : 10.1002/ece3.258

C. E. Carver, A. L. Mallet, and B. Vercaemer, Biological synopsis of the solitary tunicate Ciona intestinalis, Can. Man. Rep. Fish. Aquat. Sci, pp.2746-55, 2006.

M. L. Nydam and R. G. Harrison, INTROGRESSION DESPITE SUBSTANTIAL DIVERGENCE IN A BROADCAST SPAWNING MARINE INVERTEBRATE, Evolution, vol.59, issue.2, pp.429-442, 2011.
DOI : 10.1111/j.1558-5646.2010.01153.x

S. Bouchemousse, C. Haag-liautard, N. Bierne, and F. Viard, Past and contemporary introgression between two strongly differentiated Ciona species as revealed by a post-genomic SNPs panel, pp.10-1101, 2015.

M. Rius, X. Turon, G. Bernardi, F. A. Volckaert, and F. Viard, Marine invasion genetics: from spatio-temporal patterns to evolutionary outcomes, Biological Invasions, vol.2, issue.3, pp.869-885, 2015.
DOI : 10.1007/s10530-014-0792-0
URL : https://hal.archives-ouvertes.fr/hal-01218812

M. Rius, Range expansions across ecoregions: interactions of climate change, physiology and genetic diversity, Global Ecology and Biogeography, vol.13, issue.1, pp.76-88, 2014.
DOI : 10.1111/geb.12105

O. Affinito, sp. A, Marine Ecology, vol.2, issue.4, pp.1230-1243, 2016.
DOI : 10.1111/maec.12226

G. Lambert, Ecology and natural history of the protochordates, Canadian Journal of Zoology, vol.83, issue.1, pp.34-50, 2005.
DOI : 10.1139/z04-156

A. J. Bohonak, Dispersal, Gene Flow, and Population Structure, The Quarterly Review of Biology, vol.74, issue.1, pp.21-45, 1999.
DOI : 10.1086/392950

B. P. Kinlan and S. D. Gaines, PROPAGULE DISPERSAL IN MARINE AND TERRESTRIAL ENVIRONMENTS: A COMMUNITY PERSPECTIVE, Ecology, vol.84, issue.8, pp.2007-2020, 2003.
DOI : 10.1007/BF00346739

S. E. Lester, B. I. Ruttenberg, S. D. Gaines, and B. P. Kinlan, The relationship between dispersal ability and geographic range size, Ecology Letters, vol.230, issue.8, pp.745-758, 2007.
DOI : 10.1007/BF00541659

D. J. Marshall, P. J. Krug, E. K. Kupriyanova, M. Byrne, and R. B. Emlet, The Biogeography of Marine Invertebrate Life Histories, Annual Review of Ecology, Evolution, and Systematics, vol.43, issue.1, pp.97-114, 2012.
DOI : 10.1146/annurev-ecolsys-102710-145004

K. A. Selkoe and R. J. Toonen, Marine connectivity: a new look at pelagic larval duration and genetic metrics of dispersal, Marine Ecology Progress Series, vol.436, pp.291-305, 2011.
DOI : 10.3354/meps09238

J. P. Wares, S. D. Gaines, and C. W. Cunningham, A COMPARATIVE STUDY OF ASYMMETRIC MIGRATION EVENTS ACROSS A MARINE BIOGEOGRAPHIC BOUNDARY, Evolution, vol.1991, issue.2, pp.295-306, 2001.
DOI : 10.1046/j.1365-2540.1999.00496.x

P. A. Haye, Phylogeographic Structure in Benthic Marine Invertebrates of the Southeast Pacific Coast of Chile with Differing Dispersal Potential, PLoS ONE, vol.20, issue.2, 2014.
DOI : 10.1371/journal.pone.0088613.t007

N. Galtier, B. Nabholz, S. Glemin, and G. D. Hurst, Mitochondrial DNA as a marker of molecular diversity: a reappraisal, Molecular Ecology, vol.57, issue.22, pp.4541-4550, 2009.
DOI : 10.1111/j.1365-294X.2009.04380.x

J. P. Wares and C. W. Cunningham, PHYLOGEOGRAPHY AND HISTORICAL ECOLOGY OF THE NORTH ATLANTIC INTERTIDAL, Evolution, vol.96, issue.12, pp.2455-2469, 2001.
DOI : 10.1098/rspb.2000.1028

G. J. Vermeij, From Europe to America: Pliocene to Recent trans-Atlantic expansion of cold-water North Atlantic molluscs, Proceedings of the Royal Society B: Biological Sciences, vol.5, issue.12, pp.2545-25503177, 2005.
DOI : 10.1126/science.253.5024.1099

P. B. Marko, The ???Expansion-Contraction??? model of Pleistocene biogeography: rocky shores suffer a sea change?, Molecular Ecology, vol.23, issue.1, pp.24875-24885, 2010.
DOI : 10.1111/j.1365-294X.2009.04417.x

M. N. Dawson, Phylogeography in coastal marine animals: a solution from California?, Journal of Biogeography, vol.52, issue.6, pp.723-736, 2001.
DOI : 10.1046/j.1365-2699.2001.00572.x

P. A. Hohenlohe, Limits to gene flow in marine animals with planktonic larvae: models of Littorina species around Point Conception, California, Biological Journal of the Linnean Society, vol.82, issue.2, pp.169-187, 2004.
DOI : 10.1111/j.1095-8312.2004.00318.x

S. J. Goldstien, Global Phylogeography of the Widely Introduced North West Pacific Ascidian Styela clava, PLoS ONE, vol.234, issue.2, p.16755, 2011.
DOI : 10.1371/journal.pone.0016755.s001
URL : https://hal.archives-ouvertes.fr/hal-01218838

D. G. Bock, H. J. Macisaac, and M. E. Cristescu, Multilocus genetic analyses differentiate between widespread and spatially restricted cryptic species in a model ascidian, Proceedings of the Royal Society B: Biological Sciences, vol.17, issue.6, pp.2377-2385, 2012.
DOI : 10.1016/j.tplants.2008.03.004

J. T. Carlton and G. M. Ruiz, In Biological invasions in changing ecosystems:Vectors, ecological impacts, management and predictions (ed. João Canning-Clode), pp.24-36, 2015.

M. Rius and J. A. Darling, How important is intraspecific genetic admixture to the success of colonising populations?, Trends in Ecology & Evolution, vol.29, issue.4, pp.233-242, 2014.
DOI : 10.1016/j.tree.2014.02.003

J. Roman, Diluting the founder effect: cryptic invasions expand a marine invader's range, Proceedings of the Royal Society B: Biological Sciences, vol.10, issue.4, pp.2453-24593597, 2006.
DOI : 10.1073/pnas.0501754102

A. Brante, M. Fernandez, and F. Viard, Phylogeography and Biogeography Concordance in the Marine Gastropod Crepipatella dilatata (Calyptraeidae) along the Southeastern Pacific Coast, Journal of Heredity, vol.103, issue.5, pp.630-637, 2012.
DOI : 10.1093/jhered/ess030
URL : https://hal.archives-ouvertes.fr/hal-01218836

L. Dupont, F. Viard, P. David, and J. D. Bishop, Combined effects of bottlenecks and selfing in populations of Corella eumyota, a recently introduced sea squirt in the English Channel, Diversity and Distributions, vol.25, issue.1, pp.808-817, 2007.
DOI : 10.1111/j.1472-4642.2007.00405.x

R. Collin, P. Farrell, and S. Cragg, Confirmation of the identification and establishment of the South American slipper limpet Crepipatella dilatata (Lamark 1822) (Caenogastropoda: Calyptraeidae) in Northern Spain, Aquatic Invasions, vol.4, issue.2, pp.377-380, 2009.
DOI : 10.3391/ai.2009.4.2.13

J. C. Castilla, Down under the southeastern Pacific: marine non-indigenous species in Chile, Biological Invasions, vol.3, issue.3, pp.213-232, 2005.
DOI : 10.1007/s10530-004-0198-5

A. Geoffroy, L. Gall, L. Destombe, and C. , Cryptic introduction of the red alga Polysiphonia morrowii Harvey (Rhodomelaceae, Rhodophyta) in the North Atlantic Ocean highlighted by a DNA barcoding approach, Aquatic Botany, vol.100, pp.67-71, 2012.
DOI : 10.1016/j.aquabot.2012.03.002

J. D. Bishop, The Southern Hemisphere ascidian Asterocarpa humilis is unrecognised but widely established in NW France and Great Britain, Biological Invasions, vol.84, issue.2, pp.253-260, 2013.
DOI : 10.1007/s10530-012-0286-x
URL : https://hal.archives-ouvertes.fr/hal-01218820

C. A. Maggs, EVALUATING SIGNATURES OF GLACIAL REFUGIA FOR NORTH ATLANTIC BENTHIC MARINE TAXA, Ecology, vol.39, issue.sp11, pp.108-122, 2008.
DOI : 10.1111/j.1529-8817.2006.00292.x

L. N. Cox, N. I. Zaslavskaya, and P. B. Marko, Phylogeography and trans-Pacific divergence of the rocky shore gastropod Nucella lima, J. Biogeogr, vol.41, pp.615-627, 2014.

E. Waltari and M. J. Hickerson, Late Pleistocene species distribution modelling of North Atlantic intertidal invertebrates, Journal of Biogeography, vol.75, issue.2, pp.249-260, 2013.
DOI : 10.1111/j.1365-2699.2012.02782.x

C. K. Tepolt, Adaptation in marine invasion: a genetic perspective, Biological Invasions, vol.215, issue.3, pp.887-903, 2015.
DOI : 10.1007/s10530-014-0825-8

T. Lee and S. Shin, Morphological and molecular identification of an introduced alien sea squirt (Tunicata: Ascidiacea) in Korea, Proceedings of the Biological Society of Washington, vol.127, issue.1, pp.284-297, 2014.
DOI : 10.2988/0006-324X-127.1.284

A. M. Blakeslee, J. E. Byers, and M. P. Lesser, 's North American origin, Molecular Ecology, vol.105, issue.16, pp.3684-3696, 2008.
DOI : 10.1111/j.1365-294X.2008.03865.x

J. I. Provan, R. A. Wattier, and C. A. Maggs, Phylogeographic analysis of the red seaweed Palmaria palmata reveals a Pleistocene marine glacial refugium in the English Channel, Molecular Ecology, vol.8, issue.3, pp.793-803, 2005.
DOI : 10.1111/j.1365-294X.2005.02447.x

J. Provan and K. D. Bennett, Phylogeographic insights into cryptic glacial refugia, Trends in Ecology & Evolution, vol.23, issue.10, pp.564-571010, 2008.
DOI : 10.1016/j.tree.2008.06.010

M. Panova, Glacial History of the North Atlantic Marine Snail, Littorina saxatilis, Inferred from Distribution of Mitochondrial DNA Lineages, PLoS ONE, vol.11, issue.3, 2011.
DOI : 10.1371/journal.pone.0017511.s006

R. Hartmeyer, In The Danish Ingolf Expedition, Copenhagen, vol.2, issue.7, p.275, 1924.

W. G. Van-name, The North and South American ascidians, B. Am. Mus. Nat. Hist, vol.84, pp.471-431, 1945.

N. J. Berrill, The Tunicata with an account of the Bristish species, 1950.

K. Sanamyan, N. Ciona-fleming-shenkar, A. Gittenberger, G. Lambert, M. Rius et al., Accessed through: World Register of Marine Species at http, Ascidiacea World Database, 1822.

B. I. Dybern, The Life Cycle of Ciona intestinalis (L.) f. typica in Relation to the Environmental Temperature, Oikos, vol.16, issue.1/2, pp.109-131, 1965.
DOI : 10.2307/3564870

A. Ramsay, J. Davidson, D. Bourque, and H. Stryhn, Recruitment patterns and population development of the invasive ascidian Ciona intestinalis in Prince Edward Island, Canada, Aquatic Invasions, vol.4, issue.1, pp.169-176, 2009.
DOI : 10.3391/ai.2009.4.1.17

A. Ramsay, J. Davidson, T. Landry, and G. Arsenault, Process of invasiveness among exotic tunicates in Prince Edward Island, Canada, Biological Invasions, vol.286, issue.8, pp.1311-1316, 2008.
DOI : 10.1007/s10530-007-9205-y

P. S. Sargent, T. Wells, K. Matheson, C. H. Mckenzie, and D. Deibel, First record of vase tunicate, Ciona intestinalis (Linnaeus, 1767) in coastal Newfoundland waters, BioInvasions Records, vol.2, issue.2, pp.89-98, 2013.
DOI : 10.3391/bir.2013.2.2.01

F. Iannelli, G. Pesole, P. Sordino, and C. Gissi, Mitogenomics reveals two cryptic species in Ciona intestinalis, Trends in Genetics, vol.23, issue.9, pp.419-422, 2007.
DOI : 10.1016/j.tig.2007.07.001

T. A. Hall, BioEdit: a user friendly biological sequence alignment editor and analysis program for windows 95/98/NT, Nucleic Acids Sympos. Ser, vol.41, pp.95-98, 1999.

J. Rozas, J. C. Sanchez-delbarrio, X. Messeguer, R. Rozas, and . Dnasp, DnaSP, DNA polymorphism analyses by the coalescent and other methods, Bioinformatics, vol.19, issue.18, pp.2496-2497, 2003.
DOI : 10.1093/bioinformatics/btg359

S. T. Kalinowski, hp-rare 1.0: a computer program for performing rarefaction on measures of allelic richness, Molecular Ecology Notes, vol.33, issue.1, pp.187-189, 2005.
DOI : 10.1046/j.1365-294X.2002.01612.x

L. Excoffier and H. E. Lischer, Arlequin suite ver 3.5: a new series of programs to perform population genetics analyses under Linux and Windows, Molecular Ecology Resources, vol.127, issue.3, pp.564-567, 2010.
DOI : 10.1111/j.1755-0998.2010.02847.x

K. R. Clarke and R. N. Gorley, Primer v, 2006.

M. Hasegawa, H. Kishino, and T. A. Yano, Dating of the human-ape splitting by a molecular clock of mitochondrial DNA, Journal of Molecular Evolution, vol.275, issue.3, pp.160-174, 1985.
DOI : 10.1007/BF02101694

J. Hey and R. Nielsen, Integration within the Felsenstein equation for improved Markov chain Monte Carlo methods in population genetics, Proc. Natl. Acad. Sci. USA, pp.2785-2790, 2007.
DOI : 10.1073/pnas.0611164104