S. Altschul, W. Gish, and W. Miller, Basic local alignment search tool, Journal of Molecular Biology, vol.215, issue.3, pp.403-413, 1990.
DOI : 10.1016/S0022-2836(05)80360-2

A. Baco-taylor, Food-web structure, succession, and phylogenetics on deep-sea whale skeletons, 2002.

G. Bartoli, M. Sarnthein, M. Weinelt, H. Erlenkeuser, D. Garbe-schönberg et al., Final closure of Panama and the onset of northern hemisphere glaciation, Earth and Planetary Science Letters, vol.237, issue.1-2, pp.33-44, 2005.
DOI : 10.1016/j.epsl.2005.06.020

. Brinkmeyer-r, K. Knittel, and J. Jurgens, Diversity and Structure of Bacterial Communities in Arctic versus Antarctic Pack Ice, Applied and Environmental Microbiology, vol.69, issue.11, pp.6610-6619, 2003.
DOI : 10.1128/AEM.69.11.6610-6619.2003

E. Choi, H. Kwon, and H. Koh, Colwellia asteriadis sp. nov., a marine bacterium isolated from the starfish Asterias amurensis, INTERNATIONAL JOURNAL OF SYSTEMATIC AND EVOLUTIONARY MICROBIOLOGY, vol.60, issue.8, pp.1952-1959, 2010.
DOI : 10.1099/ijs.0.016055-0

J. Cole, Q. Wang, and E. Cardenas, The Ribosomal Database Project: improved alignments and new tools for rRNA analysis, Nucleic Acids Research, vol.37, issue.Database, 2009.
DOI : 10.1093/nar/gkn879

D. Colgan, A. Mclauchlan, and G. Wilson, Histone H3 and U2 snRNA DNA sequences and arthropod molecular evolution, Australian Journal of Zoology, vol.46, issue.5, pp.419-456, 1998.
DOI : 10.1071/ZO98048

J. Deming, A. Reysenbach, and S. Macko, Evidence for the microbial basis of a chemoautotrophic invertebrate community at a whale fall on the deep seafloor: Bone-colonizing bacteria and invertebrate endosymbionts, Microscopy Research and Technique, vol.131, issue.2, pp.162-70, 1997.
DOI : 10.1002/(SICI)1097-0029(19970415)37:2<162::AID-JEMT4>3.0.CO;2-Q

D. Distel, D. Lane, and G. Olsen, Sulfur-oxidizing bacterial endosymbionts: analysis of phylogeny and specificity by 16S rRNA sequences., Journal of Bacteriology, vol.170, issue.6, pp.2506-2516, 1988.
DOI : 10.1128/jb.170.6.2506-2510.1988

D. Distel, H. Lee, and C. Cavanaugh, Intracellular coexistence of methano- and thioautotrophic bacteria in a hydrothermal vent mussel., Proceedings of the National Academy of Sciences, vol.92, issue.21, pp.9598-602, 1995.
DOI : 10.1073/pnas.92.21.9598

N. Dubilier, C. Bergin, and C. Lott, Symbiotic diversity in marine animals: the art of harnessing chemosynthesis, Nature Reviews Microbiology, vol.214, issue.10, pp.725-765, 2008.
DOI : 10.1264/jsme2.22.136

S. Duperron, The diversity of deep-sea mussels and their bacterial symbioses The Vent and Seep Biota, pp.137-67, 2010.

S. Duperron, S. Halary, and J. Lorion, Unexpected co-occurrence of six bacterial symbionts in the gills of the cold seep mussel Idas sp. (Bivalvia: Mytilidae), Environmental Microbiology, vol.24, issue.2, pp.433-478, 2008.
DOI : 10.1128/AEM.69.11.6785-6792.2003

S. Duperron, M. Laurent, and F. Gaill, Sulphur-oxidizing extracellular bacteria in the gills of Mytilidae associated with wood falls, FEMS Microbiology Ecology, vol.63, issue.3, pp.338-387, 2008.
DOI : 10.1111/j.1574-6941.2008.00438.x
URL : https://hal.archives-ouvertes.fr/hal-00715244

S. Duperron, T. Nadalig, and J. Caprais, Dual Symbiosis in a Bathymodiolus sp. Mussel from a Methane Seep on the Gabon Continental Margin (Southeast Atlantic): 16S rRNA Phylogeny and Distribution of the Symbionts in Gills, Applied and Environmental Microbiology, vol.71, issue.4, pp.1694-700, 2005.
DOI : 10.1128/AEM.71.4.1694-1700.2005
URL : https://hal.archives-ouvertes.fr/hal-00152530

S. Duperron, M. Sibuet, and B. Macgregor, Diversity, relative abundance and metabolic potential of bacterial endosymbionts in three Bathymodiolus mussel species from cold seeps in the Gulf of Mexico, Environmental Microbiology, vol.44, issue.6, pp.1423-1461, 2007.
DOI : 10.1360/04ye0001

H. Eilers, J. Pernthaler, and F. Glöckner, Culturability and In Situ Abundance of Pelagic Bacteria from the North Sea, Applied and Environmental Microbiology, vol.66, issue.7, pp.3044-51, 2000.
DOI : 10.1128/AEM.66.7.3044-3051.2000

B. Faure, S. Schaeffer, and C. Fisher, Species Distribution and Population Connectivity of Deep-Sea Mussels at Hydrocarbon Seeps in the Gulf of Mexico, PLOS ONE, vol.8, issue.12, p.118460, 2015.
DOI : 10.1371/journal.pone.0118460.t009

C. Fisher, J. Brooks, and J. Vodenichar, The Co-occurrence of Methanotrophic and Chemoautotrophic Sulfur-Oxidizing Bacterial Symbionts in a Deep-sea Mussel, Marine Ecology, vol.23, issue.4, pp.277-89, 1993.
DOI : 10.1111/j.1439-0485.1993.tb00001.x

O. Folmer, M. Black, and W. Hoeh, DNA primers for amplification of mitochondrial cytochrome C oxidase subunit I from metazoan invertebrates, Mol Mar Biol Biotechnol, vol.3, pp.294-303, 1994.

Y. Fujiwara, M. Kawato, and C. Noda, Extracellular and Mixotrophic Symbiosis in the Whale-Fall Mussel Adipicola pacifica: A Trend in Evolution from Extra- to Intracellular Symbiosis, PLoS ONE, vol.55, issue.7, p.11808, 2010.
DOI : 10.1371/journal.pone.0011808.t002

S. Gaudron, E. Demoyencourt, and S. Duperron, : Gametogenesis and Larval Biology, The Biological Bulletin, vol.222, issue.1, pp.6-16, 2012.
DOI : 10.1086/BBLv222n1p6

S. Goffredi, H. Yi, and Q. Zhang, Genomic versatility and functional variation between two dominant heterotrophic symbionts of deep-sea Osedax worms, The ISME Journal, vol.4, issue.4, pp.908-932, 2014.
DOI : 10.1038/ismej.2013.201

O. Gros and F. Gaill, Extracellular bacterial association in gills of " wood mussels, Cah Biol Mar, vol.48, pp.103-112, 2007.
URL : https://hal.archives-ouvertes.fr/hal-00753023

R. Gustafson, R. Turner, and R. Lutz, A new genus and five new species of mussels (Bivalvia, Mytilidae) from deep-sea sulfide/hydrocarbon seeps in the Gulf of Mexico, Malacologia, vol.40, pp.63-112, 1998.

N. Hassouna, B. Michot, and J. Bachellerie, The complete nucleotide sequence of mouse 28S rRNA gene. Implications for the process of size increase of the large subunit rRNA In higher eukaryotes, Nucleic Acids Research, vol.12, issue.8, pp.3563-83, 1984.
DOI : 10.1093/nar/12.8.3563

Y. Kim, S. Park, and B. Nam, Colwellia meonggei sp. nov., a novel gammaproteobacterium isolated from sea squirt Halocynthia roretzi, Antonie van Leeuwenhoek, vol.58, issue.6, pp.1021-1028, 2013.
DOI : 10.1007/s10482-013-0022-2

S. Laming, S. Duperron, and M. Cunha, Settled, symbiotic, then sexually mature: adaptive developmental anatomy in the deep-sea, chemosymbiotic mussel Idas modiolaeformis, Marine Biology, vol.92, issue.6, pp.1319-1352, 2014.
DOI : 10.1007/s00227-014-2421-y

S. Laming, S. Duperron, and S. Gaudron, Adapted to change: The rapid development of symbiosis in newly settled, fast-maturing chemosymbiotic mussels in the deep sea, Marine Environmental Research, vol.112, pp.100-112, 2015.
DOI : 10.1016/j.marenvres.2015.07.014

S. Laming, K. Szafranski, and C. Rodrigues, Fickle or Faithful: The Roles of Host and Environmental Context in Determining Symbiont Composition in Two Bathymodioline Mussels, PLOS ONE, vol.27, issue.12, p.144307, 2015.
DOI : 10.1371/journal.pone.0144307.s003
URL : https://hal.archives-ouvertes.fr/hal-01261328

D. Lane, Nucleic Acid Techniques in Bacterial Systematics, pp.115-75, 1991.

J. Lorion, B. Buge, and C. Cruaud, New insights into diversity and evolution of deep-sea Mytilidae (Mollusca: Bivalvia), Molecular Phylogenetics and Evolution, vol.57, issue.1, pp.71-83, 2010.
DOI : 10.1016/j.ympev.2010.05.027

J. Lorion, S. Kiel, and B. Faure, Adaptive radiation of chemosymbiotic deep-sea mussels, Proc Biol Sci, vol.280, 2013.

B. Methe, K. Nelson, and J. Deming, The psychrophilic lifestyle as revealed by the genome sequence of Colwellia psychrerythraea 34H through genomic and proteomic analyses, Proceedings of the National Academy of Sciences, vol.102, issue.31, pp.10913-10921, 2005.
DOI : 10.1073/pnas.0504766102

B. Meyer and J. Kuever, Molecular analysis of the distribution and phylogeny of dissimilatory adenosine-5'-phosphosulfate reductase-encoding genes (aprBA) among sulfur-oxidizing prokaryotes, Microbiology, vol.153, issue.10, pp.3478-98, 2007.
DOI : 10.1099/mic.0.2007/008250-0

S. Ratnasingham and P. Hebert, BOLD: The Barcode of Life Data System (www.barcodinglife.org)

J. Robidart, S. Bench, and R. Feldman, Metabolic versatility of the Riftia pachyptila endosymbiont revealed through metagenomics, Environmental Microbiology, vol.41, issue.3, pp.727-764, 2008.
DOI : 10.1038/nature05192

E. Southward, The Morphology of Bacterial Symbioses in the Gills of Mussels of the Genera Adipicola and Idas (Bivalvia: Mytilidae), Journal of Shellfish Research, vol.27, issue.1, pp.139-185, 2008.
DOI : 10.2983/0730-8000(2008)27[139:TMOBSI]2.0.CO;2

|. Bathymodiolus-sp, AB036710 Bathymodiolus platifrons AM236325 Bathymodiolus heckerae clone M6