H. Abe, Habitat factors affecting the geographic size variation of Japanese moles., Mammal Study, vol.21, issue.2, pp.71-87, 1996.
DOI : 10.3106/mammalstudy.21.71

H. Abe, Soil hardness, a factor affecting the range expansion of Mogera wogura in Japan., Mammal Study, vol.26, issue.1, pp.45-52, 2001.
DOI : 10.3106/mammalstudy.26.45

D. J. Alvarez-lao and N. Garcia, Chronological distribution of Pleistocene cold-adapted large mammal faunas in the Iberian Peninsula, Quaternary International, vol.212, issue.2, pp.120-128, 2010.
DOI : 10.1016/j.quaint.2009.02.029

R. J. Baker and R. D. Bradley, SPECIATION IN MAMMALS AND THE GENETIC SPECIES CONCEPT, Journal of Mammalogy, vol.87, issue.4, pp.643-662, 2006.
DOI : 10.1644/06-MAMM-F-038R2.1

H. J. Bandelt, P. Forster, and A. Rohl, Median-joining networks for inferring intraspecific phylogenies, Molecular Biology and Evolution, vol.16, issue.1, pp.37-48, 1999.
DOI : 10.1093/oxfordjournals.molbev.a026036

A. A. Bannikova, E. D. Zemlemerova, P. Colangelo, M. Sözen, M. Sevindik et al., An underground burst of diversity, 2015.

D. Canestrelli, D. Salvi, M. Maura, M. A. Bologna, and G. Nascetti, One Species, Three Pleistocene Evolutionary Histories: Phylogeography of the Italian Crested Newt, Triturus carnifex, PLoS ONE, vol.7, issue.7, p.41754, 2012.
DOI : 10.1371/journal.pone.0041754.t002

E. Capanna, Karyotype and cranial morphology of Talpa romana (Thomas) from terra typica, Mammalia, vol.45, pp.71-82, 1981.

O. Charrier, P. Dupont, A. Pornon, and N. Escaravage, Microsatellite Marker Analysis Reveals the Complex Phylogeographic History of Rhododendron ferrugineum (Ericaceae) in the Pyrenees, PLoS ONE, vol.44, issue.2, p.92976, 2014.
DOI : 10.1371/journal.pone.0092976.s003

V. J. Cleef-roders and V. D. Hoek-ostende, Dental morphology of Talpa europaea and Talpa occidentalis (Mammalia: Insectivora) with a discussion of fossil Talpa in the Pleistocene of Europe, Zoologische Mededelingen, vol.75, pp.51-68, 2001.

P. Colangelo, A. A. Bannikova, B. Krystufek, V. S. Lebedev, F. Annesi et al., Molecular systematics and evolutionary biogeography of the genus Talpa (Soricomorpha: Talpidae), Molecular Phylogenetics and Evolution, vol.55, issue.2, pp.372-380, 2010.
DOI : 10.1016/j.ympev.2010.01.038

P. Colangelo, G. Aloise, P. Franchini, F. Annesi, and G. Amori, Mitochondrial DNA reveals hidden diversity and an ancestral lineage of the bank vole in the Italian peninsula, Journal of Zoology, vol.281, issue.1, pp.42-52, 2012.
DOI : 10.1111/j.1469-7998.2011.00884.x

W. D. Collins, C. M. Bitz, M. L. Blackmon, G. B. Bonan, C. S. Bretherton et al., The Community Climate System Model Version 3 (CCSM3), The Community Climate System Model version 3 (CCSM3), pp.2122-2143, 2006.
DOI : 10.1175/JCLI3761.1

M. Corti and A. Loy, Morphometric divergence in southern European moles (Insect??vora, Talpidae), Bolletino di zoologia, vol.21, issue.2, pp.187-191, 1987.
DOI : 10.1111/j.1469-185X.1976.tb01063.x

A. Cyprien, L. Visset, and N. Carcaud, Evolution of vegetation landscapes during the Holocene in the central and downstream Loire basin (Western France) Vegetation History and Archaeobotany, pp.181-196, 2004.

J. F. Ducroz, V. Volobouev, and L. Granjon, An assessment of the systematics of arvicanthine rodents using mitochondrial DNA sequences: evolutionary and biogeographical implications, Journal of Mammalian Evolution, vol.8, issue.3, pp.173-206, 2001.
DOI : 10.1023/A:1012277012303

J. Elith, C. H. Graham, R. P. Anderson, M. Dudik, S. Ferrier et al., Novel methods improve prediction of species??? distributions from occurrence data, Ecography, vol.19, issue.2, pp.129-151, 2006.
DOI : 10.1111/j.2006.0906-7590.04596.x

G. N. Feliner, Southern European glacial refugia: A tale of tales, Taxon, vol.60, pp.365-372, 2011.

J. W. Ferguson, On the use of genetic divergence for identifying species, Biological Journal of the Linnean Society, vol.75, issue.4, pp.509-516, 2002.
DOI : 10.1046/j.1095-8312.2002.00042.x

R. Feuda, A. A. Bannikova, E. D. Zemlemerova, M. Febbraro, A. Loy et al., , through mitochondrial DNA phylogeography and species distribution modelling, Biological Journal of the Linnean Society, vol.114, issue.3, pp.495-512, 2015.
DOI : 10.1111/bij.12459

M. G. Filippucci, G. Nascetti, E. Capanna, and L. Bullini, Allozyme variation and systematic of European moles of the genus Talpa, 1987.

Y. Fourcade, J. O. Engler, D. Rodder, and J. Secondi, Mapping Species Distributions with MAXENT Using a Geographically Biased Sample of Presence Data: A Performance Assessment of Methods for Correcting Sampling Bias, PLoS ONE, vol.330, issue.5, p.97122, 2014.
DOI : 10.1371/journal.pone.0097122.s002

T. C. Giannini, D. S. Chapman, A. M. Saraiva, I. Alves-dos-santos, and J. C. Biesmeijer, Improving species distribution models using biotic interactions: a case study of parasites, pollinators and plants, Ecography, vol.33, issue.6, pp.649-656, 2013.
DOI : 10.1111/j.1600-0587.2012.07191.x

A. Gómez and D. H. Lunt, Refugia within Refugia: Patterns of Phylogeographic Concordance in the Iberian Peninsula, Phylogeography of southern European refugia, pp.155-188, 2007.
DOI : 10.1007/1-4020-4904-8_5

M. L. Gorman and R. D. Stone, The natural history of moles, 1990.

A. Grill, G. Amori, G. Aloise, I. Lisi, G. Tosi et al., Molecular phylogeography of European Sciurus vulgaris: refuge within refugia? Molecular Ecology, pp.2687-2699, 2009.

S. Guindon, J. F. Dufayard, V. Lefort, M. Anisimova, W. Hordijk et al., New Algorithms and Methods to Estimate Maximum-Likelihood Phylogenies: Assessing the Performance of PhyML 3.0, Systematic Biology, vol.59, issue.3, pp.307-321, 2010.
DOI : 10.1093/sysbio/syq010
URL : https://hal.archives-ouvertes.fr/lirmm-00511784

S. Guindon, F. Lethiec, P. Duroux, and O. Gascuel, PHYML Online--a web server for fast maximum likelihood-based phylogenetic inference, Nucleic Acids Research, vol.33, issue.Web Server, pp.557-559, 2005.
DOI : 10.1093/nar/gki352
URL : https://hal.archives-ouvertes.fr/lirmm-00105317

A. Hassanin, J. An, A. Ropiquet, T. T. Nguyen, and A. Couloux, Combining multiple autosomal introns for studying shallow phylogeny and taxonomy of Laurasiatherian mammals: Application to the tribe Bovini (Cetartiodactyla, Bovidae), Molecular Phylogenetics and Evolution, vol.66, issue.3, pp.766-775, 2013.
DOI : 10.1016/j.ympev.2012.11.003

H. Hasumi and S. Emori, K-1 Coupled GCM (MIROC) Description, 2004.

K. He, A. Shinohara, X. L. Jiang, and K. L. Campbel, Multilocus phylogeny of talpine moles (Talpini, Talpidae, Eulipotyphla) and its implications for systematics, Molecular Phylogenetics and Evolution, vol.70, pp.513-521, 2014.
DOI : 10.1016/j.ympev.2013.10.002

G. M. Hewitt, Genetic consequences of climatic oscillations in the Quaternary, Philosophical Transactions of the Royal Society B: Biological Sciences, vol.359, issue.1442, pp.183-195, 2004.
DOI : 10.1098/rstb.2003.1388

R. J. Hijmans, S. E. Cameron, J. L. Parra, P. G. Jones, and A. Jarvis, Very high resolution interpolated climate surfaces for global land areas, International Journal of Climatology, vol.18, issue.15, pp.1965-1978, 2005.
DOI : 10.1002/joc.1276

R. Hutterer, Order Soricomorpha (Eds) Mammal species of the world: A taxonomic and geographic reference, pp.220-311, 2005.

F. Jacquet, V. Nicolas, M. Colyn, B. Kadjo, R. Hutterer et al., complex, Soricomorpha), Zoologica Scripta, vol.20, issue.2, pp.131-148, 2014.
DOI : 10.1111/zsc.12039
URL : https://hal.archives-ouvertes.fr/hal-00999609

F. Jacquet, C. Denys, E. Verheyen, J. Bryja, R. Hutterer et al., Phylogeography and evolutionary history of the Crocidura olivieri complex (Mammalia, Soricomorpha): from a forest origin to broad ecological expansion across Africa, BMC Evolutionary Biology, vol.19, issue.1, p.71, 2015.
DOI : 10.1186/s12862-015-0344-y
URL : https://hal.archives-ouvertes.fr/hal-01151637

R. Jiménez, M. Burgos, and R. Diaz-de-la-guardia, ) from the South-East of the Iberian Peninsula. Implications on its Taxonomic Position, Caryologia, vol.50, issue.3, pp.253-258, 1984.
DOI : 10.1080/00087114.1984.10797705

G. C. Johns and J. C. Avise, A Comparative Summary of Genetic Distances in the Vertebrates from the Mitochondrial Cytochome b Gene, Molecular Biology and Evolution, vol.15, issue.11, pp.1481-1490, 1998.
DOI : 10.1093/oxfordjournals.molbev.a025875

R. Kass and A. Raftery, Bayes Factors, Journal of the American Statistical Association, vol.2, issue.430, pp.773-795, 1995.
DOI : 10.1080/01621459.1995.10476572

B. Kry?tufek, Skull variability of Talpa romana stankovici from Macedonia, Acta Theriologica, vol.32, pp.463-474, 1987.
DOI : 10.4098/AT.arch.87-33

B. Kry?tufek, The taxonomy of blind moles (Talpa caeca and T. stankovici, Insectivora, Mammalia) from south-eastern Europe, pp.1-16, 1994.

B. Kry?tufek, Skull analysis of small blind moles from Turkey and Iran, Folia Zoolgica, vol.50, pp.19-25, 2001.

B. Kry?tufek and V. Vohralík, Mammals of Turkey and Cyprus. Introduction, checklist, Insectivora, Koper: Science and Research Centre of the Republic of Slovenia, 2001.

B. Kry?tufek and P. Benda, The Caucasian mole Talpa caucasica ??? a new mammal for Iran, Mammalian Biology - Zeitschrift f??r S??ugetierkunde, vol.67, issue.2, pp.113-116, 2002.
DOI : 10.1078/1616-5047-00016

E. A. Lacey, J. L. Patton, and G. N. Cameron, Life Underground: The Biology Of Subterranean Rodents, Australian Mammalogy, vol.23, issue.1, 2000.
DOI : 10.1071/AM01075_BR

P. L. Leberg, Estimating allelic richness: Effects of sample size and bottlenecks, Molecular Ecology, vol.10, issue.11, pp.2445-2449, 2002.
DOI : 10.1046/j.1365-294X.2002.01612.x

P. Librado and J. Rozas, DnaSP v5: a software for comprehensive analysis of DNA polymorphism data, Bioinformatics, vol.25, issue.11, pp.1451-1452, 2009.
DOI : 10.1093/bioinformatics/btp187

A. Loy and M. Corti, (Mammalia, Insectivora, Talpidae) in Europe: A biogeographic hypothesis based on morphometric data, Italian Journal of Zoology, vol.207, issue.3, pp.277-284, 1996.
DOI : 10.1111/j.1365-2907.1982.tb00007.x

A. Loy and E. Capanna, A parapatric contact area between two species of moles (genus Talpa): character displacement investigated through the geometric morphometric of skull, Acta Zoologica Academiae Scientiarum Hungaricae, vol.44, pp.151-164, 1998.

A. Loy, M. Capula, A. Palombi, and E. Capanna, Genetic and morphometric evidence of introgression between two species of moles (Insectivora: Talpa europaea and Talpa romana) in central Italy, Journal of Zoology, vol.72, issue.3, pp.229-238, 2001.
DOI : 10.1086/282771

A. Loy, P. Colangelo, F. Annesi, and E. Capanna, Origin and evolution of Western European moles (genus Talpa, Insectivora): a multidisciplinary approach, Mammal Study, vol.30, issue.sp1, pp.13-17, 2005.
DOI : 10.3106/1348-6160(2005)30[S13:OAEOWE]2.0.CO;2

A. E. Marcy, S. Fendorf, J. L. Patton, and E. A. Hadly, Morphological Adaptations for Digging and Climate-Impacted Soil Properties Define Pocket Gopher (Thomomys spp.) Distributions, PLoS ONE, vol.24, issue.5, p.64935, 2013.
DOI : 10.1371/journal.pone.0064935.s003

I. Martínez-solano, J. Teixeira, D. Buckley, and M. García-parís, Mitochondrial DNA phylogeography of Lissotriton boscai (Caudata, Salamandridae): evidence for old, multiple refugia in an Iberian endemic, Molecular Ecology, vol.98, issue.11, pp.3375-3388, 2006.
DOI : 10.1111/j.1365-294X.2006.03013.x

F. Médail and K. Diadema, Glacial refugia influence plant diversity patterns in the Mediterranean Basin, Journal of Biogeography, vol.29, issue.7, pp.1333-1345, 2009.
DOI : 10.1111/j.1365-2699.2008.02051.x

B. Mila, Y. Surget-groba, B. Heulin, A. Gosa, and P. S. Fitze, Multilocus phylogeography of the common lizard Zootoca vivipara at the Ibero-Pyrenean suture zone reveals lowland barriers and high-elevation introgression, BMC Evolutionary Biology, vol.13, issue.1, 0192.
DOI : 10.1007/s12686-011-9548-7
URL : https://hal.archives-ouvertes.fr/hal-00931529

G. S. Miller, Catalogue of the mammals of Western Europe (Europe exclusive of Russia) in the collection of the British Museum, Trustees of the British Museum, 1912.

A. Miraldo, G. M. Hewitt, O. S. Paulo, and B. C. Emerson, Phylogeography and demographic history of Lacerta lepida in the Iberian Peninsula: multiple refugia, range expansions and secondary contact zones, BMC Evolutionary Biology, vol.42, issue.1, 2011.
DOI : 10.1111/j.1439-0469.2004.00257.x

A. J. Mitchell-jones, G. Amori, W. Bogdanowicz, B. Krystufek, P. J. Reijnders et al., The atlas of European mammals, T & AD Poyser Natural History, 1999.

P. Nehlig, Géologie du bassin de la Loire, Geosciences, vol.12, pp.10-23, 2010.

J. Niethammer and F. Krapp, Handbuch der Säugetiere Europas, 3/I: Insektenfresser, Primaten, 1990.

J. A. Nylander, MrModeltest v2. Uppsala, Sweden: Program distributed by the author, Evolutionary Biology Centre, 2004.

P. B. Pearman, A. Guisan, O. Broennimann, and C. F. Randin, Niche dynamics in space and time, Trends in Ecology & Evolution, vol.23, issue.3, pp.149-158, 2008.
DOI : 10.1016/j.tree.2007.11.005

B. M. Petrov, Taxonomy and distribution of moles (genus Talpa, Mammalia) in Macedonia, Acta Musei Macedonici Scientiarum Naturalium, vol.12, pp.117-138, 1971.

S. J. Phillips, R. P. Anderson, and R. E. Schapire, Maximum entropy modeling of species geographic distributions, Ecological Modelling, vol.190, issue.3-4, pp.231-239, 2006.
DOI : 10.1016/j.ecolmodel.2005.03.026

S. J. Phillips and M. Dudík, Modeling of species distributions with Maxent: new extensions and a comprehensive evaluation, Ecography, vol.7, issue.0, pp.161-175, 2008.
DOI : 10.1002/1097-0258(20000715)19:13<1771::AID-SIM485>3.0.CO;2-P

D. Posada and K. A. Crandall, Intraspecific gene genealogies: trees grafting into networks, Trends in Ecology & Evolution, vol.16, issue.1, pp.37-45, 2001.
DOI : 10.1016/S0169-5347(00)02026-7

J. Proust, M. Renault, P. Guennoc, and I. Thinon, Sedimentary architecture of the Loire River drowned valleys of the French Atlantic shelf, Bulletin de la Societe Geologique de France, vol.181, issue.2, pp.129-149, 2010.
DOI : 10.2113/gssgfbull.181.2.129
URL : https://hal.archives-ouvertes.fr/hal-00517012

F. Ronquist, M. Teslenko, P. Van-der-mark, D. L. Ayres, A. Darling et al., MrBayes 3.2: Efficient Bayesian Phylogenetic Inference and Model Choice Across a Large Model Space, Systematic Biology, vol.61, issue.3, pp.539-542, 2012.
DOI : 10.1093/sysbio/sys029

S. J. Sinclair, M. D. White, and G. R. Newell, How useful are species distribution models for managing biodiversity under future climates? Ecology and Society 15, 2010.

D. Swofford, PAUP: phylogenetic analysis using parsimony, 2000.

M. Stephens, N. J. Smith, and P. Donnelly, A New Statistical Method for Haplotype Reconstruction from Population Data, The American Journal of Human Genetics, vol.68, issue.4, pp.978-989, 2001.
DOI : 10.1086/319501

M. Stephens and P. Scheet, Accounting for Decay of Linkage Disequilibrium in Haplotype Inference and Missing-Data Imputation, The American Journal of Human Genetics, vol.76, issue.3, pp.449-462, 2005.
DOI : 10.1086/428594

C. S. Thaeler, An analysis of the distribution of pocket gopher species in northeastern California (Genus Thomomys), 1968.

C. Tougard, E. Renvoise, A. Petitjean, and J. P. Quere, New Insight into the Colonization Processes of Common Voles: Inferences from Molecular and Fossil Evidence, PLoS ONE, vol.1143, issue.10, p.3532, 2008.
DOI : 10.1371/journal.pone.0003532.s006
URL : https://hal.archives-ouvertes.fr/halsde-00336404

G. A. Tryfonopoulos, E. G. Thanou, S. E. Fraguedakis-tsolis, and B. P. Chondropoulos, (Mammalia, Talpidae), Journal of Zoological Systematics and Evolutionary Research, vol.92, issue.2, pp.188-193, 2009.
DOI : 10.1111/j.1439-0469.2009.00533.x

S. Varela, M. S. Lima-ribeiro, and L. C. Terribile, A Short Guide to the Climatic Variables of the Last Glacial Maximum for Biogeographers, PLOS ONE, vol.20, issue.2???3, pp.129037-129065, 2015.
DOI : 10.1371/journal.pone.0129037.s005

D. L. Warren, R. E. Glor, and M. Turelli, ENMTools: a toolbox for comparative studies of environmental niche models, Ecography, vol.62, pp.607-611, 2010.
DOI : 10.1111/j.1600-0587.2009.06142.x

G. R. Witte, Der Maulwurf Talpa europaea, 1997.