A. D. Anbar, Proterozoic Ocean Chemistry and Evolution: A Bioinorganic Bridge?, Science, vol.297, issue.5584, p.601, 2002.
DOI : 10.1126/science.1069651

URL : http://citeseerx.ist.psu.edu/viewdoc/summary?doi=

R. A. Andersen and J. C. Bailey, PHYLOGENETIC ANALYSIS OF 32 STRAINS OF VAUCHERIA (XANTHOPHYCEAE) USING THE rbcL GENE AND ITS TWO FLANKING SPACER REGIONS1, Journal of Phycology, vol.28, issue.suppl., pp.583-604, 2002.
DOI : 10.1046/j.1529-8817.2002.01144.x

A. Arber, Water plants. a study of aquatic angiosperms, p.609, 1920.

J. M. Archibald, The Puzzle of Plastid Evolution, Current Biology, vol.19, issue.2, pp.81-89, 2009.
DOI : 10.1016/j.cub.2008.11.067

M. J. Attrill, A Rehabilitated Estuarine Ecosystem, p.611, 1998.
DOI : 10.1007/978-1-4419-8708-2

C. F. Baer, M. M. Miyamoto, and D. R. Denver, Mutation rate variation in multicellular eukaryotes: causes and consequences, Nature Reviews Genetics, vol.2, issue.8, pp.619-631, 2007.
DOI : 10.1038/nrg2158

J. C. Bailey, Phaeothamniophyceae. The Tree of Life Web Project Available at, 2010.

C. Nova, A new Lineage of Chromophytes based upon photosynthetic pigments, rbcL 617 sequence analysis and ultrastructure, Protist, vol.149, pp.245-63

I. Bartsch and R. Kuhlenkamp, The marine macroalgae of Helgoland (North Sea): an annotated list of records between 1845 and 1999, Helgoland Marine Research, vol.54, issue.4, pp.160-189, 2000.
DOI : 10.1007/s101520000050

B. Becker and B. Marin, Streptophyte algae and the origin of embryophytes, Annals of Botany, vol.103, issue.7, pp.999-1004, 2009.
DOI : 10.1093/aob/mcp044

M. J. Beilby, Salt tolerance at single cell level in giant-celled Characeae. Front, 2015.
DOI : 10.3389/fpls.2015.00226

URL : http://doi.org/10.3389/fpls.2015.00226

J. H. Belcher, C. F. Carter, and A. F. Peters, An unknown filamentous brown alga 625 (Phaeophyceae) from a domestic freshwater aquarium in Northampton UK, p.13, 2009.

L. Bergstrom, A. Tatarenkov, K. Johannesson, R. B. Jonsson, and L. Kautsky, Genetic 628, 2005.

C. E. Blank, Phylogenetic distribution of compatible solute synthesis genes support a 631 freshwater origin for cyanobacteria, J. Phycol, vol.49, pp.880-895

C. E. Blank, Origin and early evolution of photosynthetic eukaryotes in freshwater environments: reinterpreting proterozoic paleobiology and biogeochemical processes in light of trait evolution, Journal of Phycology, vol.16, issue.197, pp.1040-1055
DOI : 10.1111/jpy.12111

A. J. Brook, The biology of desmids, p.636, 1981.

J. Brown and U. Sorhannus, A molecular genetic timescale for the diversification of 637, 2010.

R. V. Burne, J. Bauld, and P. De-deckker, Saline lake charophytes and their geological 640 significance, J. Sediment. Res, vol.50, pp.281-293, 1980.

G. G. Lilitska and O. P. Bilous, Ulvophyceae. Algae of Ukraine: diversity, 643 nomenclature, taxonomy, ecology and geography, Chlorophyta. Gantner Verlag, vol.3, issue.644, pp.20-61, 2011.

C. Vega, G. Wiens, and J. J. , Why are there so few fish in the sea?, Proc. Biol. Sci, p.649, 2012.

T. Cavalier-smith, Megaphylogeny, Cell Body Plans, Adaptive Zones: Causes and Timing of Eukaryote Basal Radiations, Journal of Eukaryotic Microbiology, vol.7, issue.1, pp.26-33, 2009.
DOI : 10.1111/j.1550-7408.2008.00373.x

E. Cocquyt, H. Verbruggen, F. Leliaert, and O. De-clerck, Evolution and Cytological Diversification of the Green Seaweeds (Ulvophyceae), Molecular Biology and Evolution, vol.27, issue.9, pp.2052-61, 2010.
DOI : 10.1093/molbev/msq091

J. Collén, Genome structure and metabolic features in the red seaweed Chondrus 658 crispus shed light on evolution of the Archaeplastida, Proc. Natl. Acad. Sci. U. S. A, vol.659, issue.110, pp.5247-52, 2013.

S. M. Colman, E. B. Karabanov, and C. H. Nelson, Quaternary sedimentation and 661 subsidence history of Lake Baikal, Siberia, based on seismic stratigraphy and coring, J, p.662, 2003.

. Coleochaetophyceae, Handbook of the Protists, p.665

T. Dagan, Genomes of Stigonematalean Cyanobacteria (Subsection V) and the Evolution of Oxygenic Photosynthesis from Prokaryotes to Plastids, Genome Biology and Evolution, vol.5, issue.1, pp.31-668, 2013.
DOI : 10.1093/gbe/evs117

J. Davis and Y. Lipkin, Lamprothamnium prosperity in permanently hypersaline water, p.670, 1986.
DOI : 10.1007/bf02560200

C. J. Dawes, Marine Botany, p.672, 1998.

T. R. Deason, Filamentous and Colonial Soil Algae from Dauphin Island, Alabama, Transactions of the American Microscopical Society, vol.88, issue.2, p.673, 1969.
DOI : 10.2307/3224496

S. M. Dittami, A. Gravot, S. Goulitquer, S. Rousvoal, A. F. Peters et al., Towards deciphering dynamic changes and evolutionary mechanisms 676 involved in the adaptation to low salinities in Ectocarpus (brown algae), Plant J, vol.71, pp.366-377

M. C. Feller, Human impacts on streams and rivers, p.678, 2009.

T. Friedl and C. J. Kelly, Phylogenetic relationships of green algae assigned to the 680, 2002.

L. Gall, L. Saunders, and G. W. , A nuclear phylogeny of the Florideophyceae (Rhodophyta) inferred from combined EF2, small subunit and large subunit ribosomal DNA: Establishing the new red algal subclass Corallinophycidae, Molecular Phylogenetics and Evolution, vol.43, issue.3, pp.1118-1148, 2007.
DOI : 10.1016/j.ympev.2006.11.012

U. Geissler, Die salzbelastete Flusstrecke der Werra -ein Binnenlandstandort für 689, 1983.

R. J. Gibbs, Mechanisms Controlling World Water Chemistry, Science, vol.170, issue.3962, pp.1088-90, 1970.
DOI : 10.1126/science.170.3962.1088

E. Goldberg, Minor elements in sea water, p.692, 1965.

. Oceanography, Academic Press, pp.163-196

J. M. Graham, J. A. Kranzfelder, and M. T. Auer, Light and temperature as factors 694 regulating seasonal growth and distribution of Ulothrix zonata (Ulvophyceae), J. Phycol, vol.695, issue.21, pp.228-234, 2004.

M. D. Guiry and G. M. Guiry, AlgaeBase. World-wide electronic publication Available 697 at, 2015.
DOI : 10.1111/j.0022-3646.2003.03906001_56.x

P. W. Haines, M. Hand, and M. Sandiford, Palaeozoic synorogenic sedimentation in central and northern Australia: A review of distribution and timing with implications for the evolution of intracontinental orogens, Australian Journal of Earth Sciences, vol.133, issue.41, pp.911-928, 2001.
DOI : 10.1046/j.1440-0952.2001.00909.x

J. Hall and R. M. Mccourt, Conjugating Green Algae Including Desmids, p.702, 2015.
DOI : 10.1016/B978-0-12-385876-4.00009-8

T. U. Hammer, Saline lake ecosystems of the world, p.705, 1986.

T. Han and B. Runnegar, Megascopic eukaryotic algae from the 2.1-billion-year-old 707, 1992.

DOI : 10.1046/j.1529-8817.2002.01151.x

F. Hauck, Verzeichniss der im Golfe von Triest gesammelten Meeralgen. (Fortsetzung), 1876.
DOI : 10.1007/bf01622805

R. Hecky and P. Kilham, Nutrient limitation of phytoplankton in freshwater and 717 marine environments: A review of recent evidence on the effects of enrichment, Limnol, p.718, 1988.

. Chlorophyta, of the Svalbard Archipelago: diversity, biogeography and, p.721

D. O. Hessen, Solar radiation and the evolution of life, p.723, 2008.

M. Hirano, Freshwater algae in the Antartic regions, 1965.

L. Hoffmann, Algae of terrestrial habitats, The Botanical Review, vol.10, issue.26, pp.77-105, 1989.
DOI : 10.1007/BF02858529

R. Horimoto, Y. Masakiyo, K. Ichihara, and S. Shimada, Enteromorpha-like Ulva 729 (Ulvophyeae, Chlorophyta) growing in the Todoroki River, Ishigaki Island, Japan, with 730 species reference to Ulva meridionalis Horimoto et Shimada sp, nov. Bull. Natl. Sci. Mus, p.731, 2011.

D. L. Huston, S. Pehrsson, B. M. Eglington, and K. Zaw, The Geology and Metallogeny of Volcanic-Hosted Massive Sulfide Deposits: Variations through Geologic Time and with Tectonic Setting, Economic Geology, vol.105, issue.3, pp.571-591, 2010.
DOI : 10.2113/gsecongeo.105.3.571

K. Ichihara, S. Arai, and S. Shimada, cDNA cloning of a lectin-like gene preferentially 736 expressed in freshwater from the macroalga Ulva limnetica Phycol, p.737, 2009.

K. Ichihara, F. Mineur, and S. Shimada, ISOLATION AND TEMPORAL EXPRESSION ANALYSIS OF FRESHWATER-INDUCED GENES IN ULVA LIMNETICA (ULVALES, CHLOROPHYTA)1, Journal of Phycology, vol.20, issue.3, pp.584-590, 2011.
DOI : 10.1111/j.1529-8817.2011.01001.x

C. Jackson and A. Reyes-prieto, The mitochondrial genomes of the glaucophytes 749, 2014.

. Gloeochaete-wittrockiana, Cyanoptyche gloeocystis: multilocus phylogenetics suggests a 750 monophyletic archaeplastida, Genome Biol. Evol, vol.6, pp.2774-85

D. John and F. Rindi, Filamentous (Nonconjugating) and??Plantlike Green Algae, p.752, 2015.
DOI : 10.1016/B978-0-12-385876-4.00008-6

J. D. Wehr, R. Sheath, and J. P. Kociolek, Freshwater algae of North America: 753 ecology and classification, pp.375-754

D. T. Johnston, F. Wolfe-simon, A. Pearson, and A. H. Knoll, Anoxygenic 755 photosynthesis modulated Proterozoic oxygen and sustained Earth's middle age, Proc. Natl, p.756, 2009.
DOI : 10.1073/pnas.0909248106

URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC2753640

U. Karsten and F. Rindi, Ecophysiological performance of an urban strain of the 758 aeroterrestrial green alga Klebsormidium sp, Eur. J, p.759, 2010.

A. K. Khfaji and T. A. Norton, The effects of salinity on the distribution of Fucus ceranoides, Estuarine and Coastal Marine Science, vol.8, issue.5, pp.433-439, 1979.
DOI : 10.1016/0302-3524(79)90060-4

L. Kies and B. P. Kremer, Typification of the Glaucocystophyta, Taxon, vol.35, issue.1, p.128, 1986.
DOI : 10.2307/1221049

B. Kloareg and R. S. Quatrano, Structure of the cell-walls of marine-algae and 770 ecophysiological functions of the matrix polysaccharides, Oceanogr. Mar. Biol, vol.26, pp.259-315, 1988.

D. W. Lam, H. Verbruggen, G. W. Saunders, and M. L. Vis, Multigene phylogeny of the red algal subclass Nemaliophycidae, Molecular Phylogenetics and Evolution, vol.94, pp.730-736, 2016.
DOI : 10.1016/j.ympev.2015.10.015

S. R. Cor, on the French Atlantic coast: biology and ecology, Acta Bot. Gallica, vol.160, pp.107-775

R. K. Lee, A catalogue of the marine algae of the Canadian Arctic, Natl. Mus. Can, p.777, 1980.

O. Clerck, Phylogeny and molecular evolution of the green algae, Crit. Rev. Plant Sci. 781, vol.31, pp.1-46, 2012.

F. Leliaert, H. Verbruggen, and F. W. Zechman, Into the deep: New discoveries at the base of the green plant phylogeny, BioEssays, vol.124, issue.9, pp.683-92, 2011.
DOI : 10.1002/bies.201100035

J. M. Lopez-bautista, Molecular systematics of the subaerial green algal order 792, 2006.

W. Martin, J. Baross, D. Kelley, and M. J. Russell, Hydrothermal vents and the origin of 799 life, Nature Rev. Microbiol, vol.6, pp.805-819, 2008.

J. H. Martin, S. E. Fitzwater, and R. M. Gordon, Iron deficiency limits phytoplankton growth in Antarctic waters, Global Biogeochemical Cycles, vol.64, issue.1, pp.5-12, 1990.
DOI : 10.1029/GB004i001p00005

. Algae-bodanela, . Ectocarpus, and P. Heribaudiella, Phaeophyceae) on the basis of 804 rbcL sequences and morphological characters, Phycologia, vol.46, pp.429-439

M. F. Wojciechowski and R. W. Hoshaw, Phylogeny of the conjugating green algae 807 (Zygnemophyceae) based on rbcl sequences, J. Phycol, vol.36, pp.747-758, 2000.

G. I. Mcfadden, PRIMARY AND SECONDARY ENDOSYMBIOSIS AND THE ORIGIN OF PLASTIDS, Journal of Phycology, vol.10, issue.6, p.809, 2001.
DOI : 10.1038/22099

G. I. Mcfadden, Origin and Evolution of Plastids and Photosynthesis in Eukaryotes, Cold Spring Harbor Perspectives in Biology, vol.6, issue.4, p.811, 2014.
DOI : 10.1101/cshperspect.a016105

R. F. Mullins, A molecular phylogenetic assessment of Pseudendoclonium Available at: 818 http://scholarworks.umass, pp.1-4047, 2007.

J. R. Necchi, River Algae, p.820, 2016.
DOI : 10.1007/978-3-319-31984-1

R. Nielsen, Marine algae within calcareous shells from New Zealand, New Zealand Journal of Botany, vol.16, issue.3, p.824, 1988.
DOI : 10.1111/j.1756-1051.1986.tb00908.x

J. A. Ochoa-de-alda, R. Esteban, M. L. Diago, and J. Houmard, The plastid ancestor originated among one of the major cyanobacterial lineages, Nature Communications, vol.294, pp.4937-827, 2014.
DOI : 10.1038/ncomms5937

J. L. Olsen, The genome of the seagrass Zostera marina reveals angiosperm adaptation to the sea, Nature, vol.162, issue.7590, pp.331-335, 2016.
DOI : 10.1038/nature16548

URL : https://hal.archives-ouvertes.fr/hal-01289099

L. W. Parfrey, D. J. Lahr, A. H. Knoll, and L. A. Katz, Estimating the timing of early 831 eukaryotic diversification with multigene molecular clocks, Proc. Natl. Acad. Sci. U. S. A, vol.832, issue.108, pp.13624-13633, 2011.

D. L. Pinti, The Origin and Evolution of the Oceans, p.837, 2005.
DOI : 10.1007/10913406_4

H. Reisse and J. , Advances in Astrobiology and Biogeophysics, Lect. Astrobiol, p.838
URL : https://hal.archives-ouvertes.fr/hal-00120167

D. Moreira, An early-branching freshwater cyanobacterium at the origin of plastids, p.841, 2017.

J. A. Raven, Constraints on marine algal invasion of low? salinity environments: sex in 845, 1999.

V. Reeb and D. Bhattacharya, Red Algae in the Genomic Age, Seckbach, J. & 847, 2010.

. Origin, Life in Extreme Habitats and Astrobiology, pp.849-409

R. H. Reed, On the conspecificity of marine and freshwater Bangia in Britain, p.851, 1980.

R. H. Reed, (Rhodophyta, Bangiales): the osmotic role of floridoside, British Phycological Journal, vol.103, issue.3, pp.211-218, 1985.
DOI : 10.1080/00071618500650221

H. J. Bohnert, H. Philippe, and B. F. Lang, Monophyly of primary photosynthetic 859 eukaryotes: green plants, red algae, and glaucophytes, Curr. Biol, vol.15, pp.1325-1330, 2005.

R. Salminen, Geochemical Atlas of Europe, Part 1, Background Information, 861 Methodology and Maps, 2005.

M. V. Sanchez-puerta, P. I. Leonardi, C. J. O-'kelly, and E. J. Cáceres, BELONGS TO THE ORDER CHAETOPELTIDALES (CLASS CHLOROPHYCEAE), EVIDENCE FROM ULTRASTRUCTURE AND SSU RDNA SEQUENCE DATA, Journal of Phycology, vol.42, issue.4, pp.943-950, 2006.
DOI : 10.1111/j.1529-8817.2006.00245.x

M. J. Sanderson, J. L. Thorne, N. Wikström, and K. Bremer, Molecular evidence on plant divergence times, American Journal of Botany, vol.91, issue.10, pp.1656-65, 2004.
DOI : 10.3732/ajb.91.10.1656

D. W. Schindler, Evolution of Phosphorus Limitation in Lakes, Science, vol.195, issue.4275, pp.260-262, 1977.
DOI : 10.1126/science.195.4275.260

R. Schloesser and J. L. Blum, SPHACELARIA LACUSTRIS SP. NOV., A FRESHWATER BROWN ALGA FROM LAKE MICHIGAN, Journal of Phycology, vol.14, issue.2, pp.201-207, 1980.
DOI : 10.1111/j.1529-8817.1966.tb04595.x

H. Schubert and I. Blindow, Charophytes of the Baltic Sea, Oberreifenberg: Koeltz, vol.871, 2004.

R. W. Scotland and A. H. Wortley, How Many Species of Seed Plants Are There?, Taxon, vol.52, issue.1, p.52101, 2003.
DOI : 10.2307/3647306

R. Sheath and K. M. Cole, Distribution and salinity adaptations of Bangia 881 atropurpurea (Rhodophyta), a putative migrant into the Laurentian Great Lakes, J. Phycol, vol.882, issue.16, pp.412-420, 1980.

R. G. Sheath and J. A. Hambrook, Freshwater ecology, p.884, 1990.

R. G. Sheath and M. L. Vis, Red algae Freshwater algae of North America: ecology and classification, p.887, 2015.

V. A. Shepherd and M. J. Beilby, The effect of an extracellular mucilage on the response 889 to osmotic shock in the charophyte alga Lamprothamnium papulosum, J. Membr. Biol, vol.890, issue.170, pp.229-242, 1999.

A. R. Sherwood, T. B. Shea, and R. G. Sheath, European freshwater Hildenbrandia (Hildenbrandiales, Rhodophyta) has not been derived from multiple invasions from marine habitats, Phycologia, vol.41, issue.1, pp.87-95, 2002.
DOI : 10.2216/i0031-8884-41-1-87.1

T. Silberfeld, J. W. Leigh, H. Verbruggen, C. Cruaud, B. De-reviers et al., 895 A multi-locus time-calibrated phylogeny of the brown algae, p.896, 2010.

. Phaeophyceae, Investigating the evolutionary nature of the " brown algal crown radiation, p.897

T. Silberfeld, M. L. Racault, R. L. Fletcher, A. Couloux, F. Rousseau et al., Systematics and evolutionary history of pyrenoid-bearing taxa in brown algae 900 (Phaeophyceae), Eur. J. Phycol, vol.899, issue.46, pp.361-377, 2011.

T. Silberfeld, F. Rousseau, and B. De-reviers, An Updated Classification of Brown Algae (Ochrophyta, Phaeophyceae), Cryptogamie, Algologie, vol.35, issue.2, pp.117-156, 2014.
DOI : 10.7872/crya.v35.iss2.2014.117

P. C. Silva and M. Chacana, Marine algae from Islas San Félix y San Ambrosio (Chilean 904 Oceanic Islands), Cryptogamie Algologie, vol.26, pp.103-118, 2005.

G. R. South and A. Whittick, An Introduction to Phycology, p.909, 2009.

B. Stuercke and &. K. Mcdermid, Variation in algal turf species composition and 911 abundance on two Hawaiian shallow subtidal reefs, Cryptogamie Algologie, vol.25, pp.353-365, 2004.

A. Tatarenkov, L. Bergström, R. B. Jönsson, E. A. Serrão, L. Kautsky et al., Intriguing asexual life in marginal populations of the brown seaweed Fucus vesiculosus, Molecular Ecology, vol.30, issue.2, pp.647-51, 2005.
DOI : 10.1111/j.1365-294X.2005.02425.x

D. D. Tupa, An investigation of certain chaetophoralean algae, Nova Hedwigia Beiheft, vol.923, issue.46, pp.1-155, 1974.

G. Valet, Approche paléoécologique du monde des Dasycladales à partir de l'écologie 925, 1979.

C. De-vargas, Eukaryotic plankton diversity in the sunlit ocean, Science, vol.48, issue.1, p.3481261605, 2015.
DOI : 10.1016/j.femsec.2004.10.006

O. Vinogradova and T. Darienko, Terrestrial algae of hypersaline environments of the 929, 2008.

M. L. Vis and R. G. Sheath, Distribution and systematics of Batrachospermum (Batrachospermales, Rhodophyta) in North America. 9. Section Batrachospermum: description of five new species, Phycologia, vol.35, issue.2, pp.124-134, 1996.
DOI : 10.2216/i0031-8884-35-2-124.1

J. D. Wehr and R. G. Sheath, Freshwater habitats of algae, p.934, 2003.

J. D. Wehr and R. G. Sheath, Freshwater algae of North America: ecology and 937 classification, p.938, 2003.

J. D. Wehr, R. Stancheva, K. Truhn, and R. G. Sheath, Discovery of the rare freshwater 939 brown alga Pleurocladia lacustris (Ectocarpales, Phaeophyceae) in California Streams, Am. Nat, vol.73, pp.148-157, 2013.

C. H. Wellman and P. K. Strother, The terrestrial biota prior to the origin of land plants (embryophytes): a review of the evidence, Palaeontology, vol.4, issue.1, pp.601-627, 2015.
DOI : 10.1111/pala.12172

J. West and G. Kraft, Ectocarpus siliculosus (Dillwyn) Lyngb. from Hopkins River 944, 1996.

. Falls, Victoria -the first record of a freshwater brown alga in Australia, Muelleria, vol.9, pp.29-33

B. A. Whitton and D. M. John, Coded List of Freshwater Algae 2014, p.947, 2014.

B. Becker, Origin of land plants: Do conjugating green algae hold the key? BMC Evol, p.950, 2011.

M. J. Wynne, A checklist of benthic marine algae of the tropical and subtropical 952 western Atlantic: third revision, Nova Hedwigia Beihefte, vol.140, pp.1-144, 2011.

W. Spalding, H. Pueschel, C. M. Buchheim, J. A. Verghese, B. Hanisak et al., An 958 unrecognized ancient lineage of green plants persists in deep marine waters, J. Phycol, vol.959, issue.46, pp.1288-1295, 2010.

J. P. Zehr and B. B. Ward, Nitrogen Cycling in the Ocean: New Perspectives on Processes and Paradigms, Applied and Environmental Microbiology, vol.68, issue.3, pp.1015-1024, 2002.
DOI : 10.1128/AEM.68.3.1015-1024.2002

H. Zhu, F. Leliaert, Z. Zhao, S. Xia, Z. Hu et al., Ulvella tongshanensis (Ulvellaceae, Chlorophyta), a new freshwater species from China, and an emended morphological circumscription of the genus Ulvella, Fottea, vol.15, issue.1, pp.95-104, 2015.
DOI : 10.5507/fot.2015.008

A. Falace, F. Vita, J. C. Braga, and B. Antoli?, First freshwater coralline alga and the 967 role of local features in a major biome transition, Sci. Rep, vol.6, p.968, 2016.

. Salminen, The term brackish 974 water refers to a mixture of seawater and fresh water with salinities ranging from 0.5 to 30 975 g·L -1 . 976 Seawater Fresh water Overall salinity 30-40 g·L -1 <0, Ott Wetzel, vol.5, p.1, 1988.

. Silicate, SiO4 4-) 0-180 µM 1-500 µM