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This work was also supported by a grant from Agence Nationale de la Recherche under the LabEx ANR-10-LABX-0003-BCDiv, in the program Investissements d'avenir " n ANR-11-IDEX-0004-02. 3D reconstructions were realized in the 3D platform of the EvoFonc team of the UMR 7205 We thank Miguel Garcia-Sanz (UMS2700 CNRS) for the scans of the insects, Patricia Wills (UMS2700 CNRS) for her help during tomography reconstructions, and Anthony Herrel (UMR 7179 CNRS) for his technical help. We thank Jun-Jie Gu for making photographs of type specimens of Chinese fossil species, The authors thank the team of CeMIM (USM 0504, Department RDDM, MNHN) for access to their three-dimensional workstation. We sincerely thank G. Beccaloni (NHM), pp.16-17 ,
Rasnitsyn for their authorizations of use of photographs of habitus ,
are first authors with equal ranks ,