J. Vacelet and C. Donadey, Electron microscope study of the association between some sponges and bacteria, Journal of Experimental Marine Biology and Ecology, vol.30, issue.3, pp.301-314, 1977.
DOI : 10.1016/0022-0981(77)90038-7

U. Hentschel, Microbial Diversity of Marine Sponges, Prog Mol Subcell Biol, vol.37, pp.59-88, 2003.
DOI : 10.1007/978-3-642-55519-0_3

U. Hentschel, K. M. Usher, and M. W. Taylor, Marine sponges as microbial fermenters, FEMS Microbiology Ecology, vol.55, issue.2, pp.167-177, 2006.
DOI : 10.1111/j.1574-6941.2005.00046.x

URL : https://academic.oup.com/femsec/article-pdf/55/2/167/18100192/55-2-167.pdf

S. Schmitt, Assessing the complex sponge microbiota: core, variable and species-specific bacterial communities in marine sponges, The ISME Journal, vol.12, issue.3, pp.564-576, 2012.
DOI : 10.1007/BF02010474

R. L. Simister, P. Deines, E. S. Botté, N. S. Webster, and M. W. Taylor, Sponge-specific clusters revisited: a comprehensive phylogeny of sponge-associated microorganisms, Environmental Microbiology, vol.7, issue.2, pp.517-524, 2012.
DOI : 10.1007/BF02010474

H. M. Reiswig, Partial Carbon and Energy Budgets of the Bacteriosponge Verohgia fistularis (Porifera: Demospongiae) in Barbados, Marine Ecology, vol.130, issue.1, pp.273-293, 1981.
DOI : 10.1007/BF00387115

J. Kamke, M. W. Taylor, and S. Schmitt, Activity profiles for marine sponge-associated bacteria obtained by 16S rRNA vs 16S rRNA gene comparisons, The ISME Journal, vol.55, issue.4, pp.498-508, 2010.
DOI : 10.1038/ismej.2009.143

URL : http://www.nature.com/ismej/journal/v4/n4/pdf/ismej2009143a.pdf

E. C. Giles, Bacterial community profiles in low microbial abundance sponges, FEMS Microbiology Ecology, vol.83, issue.1, pp.232-241, 2013.
DOI : 10.1111/j.1574-6941.2012.01467.x

R. Simister, M. W. Taylor, K. M. Rogers, P. J. Schupp, and P. Deines, Temporal molecular and isotopic analysis of active bacterial communities in two New Zealand sponges, FEMS Microbiology Ecology, vol.85, issue.1, pp.10-1111, 2013.
DOI : 10.1111/1574-6941.12109

U. Hentschel, Molecular Evidence for a Uniform Microbial Community in Sponges from Different Oceans, Applied and Environmental Microbiology, vol.68, issue.9, pp.4431-4440, 2002.
DOI : 10.1128/AEM.68.9.4431-4440.2002

J. Piel, Metabolites from symbiotic bacteriaThis review is dedicated to Professor Axel Zeeck on the occasion of his 65th birthday., Natural Product Reports, vol.21, issue.4, pp.519-538, 2004.
DOI : 10.1039/b310175b

D. J. Faulkner, Highlights of marine natural products chemistry (1972???1999), Natural Product Reports, vol.17, issue.1, pp.1-6, 2000.
DOI : 10.1039/a909113k

D. Schwarzer and M. A. Marahiel, Multimodular biocatalysts for natural product assembly, Naturwissenschaften, vol.88, issue.3, pp.93-101, 2001.
DOI : 10.1007/s001140100211

T. K. Kim and J. A. Fuerst, Diversity of polyketide synthase genes from bacteria associated with the marine sponge Pseudoceratina clavata: culture-dependent and culture-independent approaches, Environmental Microbiology, vol.178, issue.8, pp.1460-1470, 2006.
DOI : 10.1073/pnas.96.21.11740

Y. Kashman, Ptilomycalin A: a novel polycyclic guanidine alkaloid of marine origin, Journal of the American Chemical Society, vol.111, issue.24, pp.8925-8926, 1989.
DOI : 10.1021/ja00206a029

E. A. Jares-erijman, A. L. Ingrum, J. R. Carney, K. L. Rinehart, and R. Sakai, Polycyclic guanidine-containing compounds from the Mediterranean sponge Crambe crambe: the structure of 13,14,15-isocrambescidin 800 and the absolute stereochemistry of the pentacyclic guanidine moieties of the crambescidins, The Journal of Organic Chemistry, vol.58, issue.18, pp.4805-4808, 1993.
DOI : 10.1021/jo00070a012

E. A. Jares-erijman, R. Sakai, and K. L. Rinehart, Crambescidins: new antiviral and cytotoxic compounds from the sponge Crambe crambe, The Journal of Organic Chemistry, vol.56, issue.19, pp.5712-5715, 1991.
DOI : 10.1021/jo00019a049

R. Tavares, Isolation of crambescidin 800 from Monanchora arbuscula (Porifera), Biochemical Systematics and Ecology, vol.22, issue.6, pp.645-646, 1994.
DOI : 10.1016/0305-1978(94)90078-7

A. D. Patil, -CD4 Dissociation, The Journal of Organic Chemistry, vol.62, issue.6, pp.1814-1819, 1997.
DOI : 10.1021/jo962084t

URL : https://hal.archives-ouvertes.fr/hal-00302201

G. C. Harbour, Ptilocaulin and isoptilocaulin, antimicrobial and cytotoxic cyclic guanidines from the Caribbean sponge Ptilocaulis aff. P. spiculifer (Lamarck, 1814), Journal of the American Chemical Society, vol.103, issue.18, pp.5604-5606, 1981.
DOI : 10.1021/ja00408a071

B. B. Snider and Z. Shi, Biomimetic synthesis of the pentacyclic nucleus of ptilomycalin A, Journal of the American Chemical Society, vol.116, issue.2, pp.549-557, 1994.
DOI : 10.1021/ja00081a015

R. G. Berlinck, Channel Blocker Activity of Crambescidin 816, Journal of Natural Products, vol.56, issue.7, pp.1007-1015, 1993.
DOI : 10.1021/np50097a004

M. A. Becerro, N. I. Lopez, X. Turon, and M. J. Uriz, Antimicrobial activity and surface bacterial film in marine sponges, Journal of Experimental Marine Biology and Ecology, vol.179, issue.2, pp.195-205, 1994.
DOI : 10.1016/0022-0981(94)90114-7

M. J. Uriz, M. A. Becerro, J. M. Tur, and X. Turon, Location of toxicity within the Mediterranean sponge Crambe crambe (Demospongiae: Poecilosclerida), Marine Biology, vol.178, issue.4, pp.583-590, 1996.
DOI : 10.1007/BF00351039

M. Sarà, Associazionia fra Poriferi e alghe in acque superficiali det litorale marino. Ricerca scient 36, pp.277-282, 1966.

V. Thiel, S. C. Neulinger, T. Staufenberger, R. Schmaljohann, and J. Imhoff, Spatial distribution of sponge-associated bacteria in the Mediterranean sponge Tethya aurantium, FEMS Microbiology Ecology, vol.59, issue.1, pp.47-63, 2007.
DOI : 10.1111/j.1574-6941.2006.00217.x

A. Blazejak, C. Erséus, R. Amann, and N. Dubilier, Coexistence of Bacterial Sulfide Oxidizers, Sulfate Reducers, and Spirochetes in a Gutless Worm (Oligochaeta) from the Peru Margin, Applied and Environmental Microbiology, vol.71, issue.3, pp.1553-1561, 2005.
DOI : 10.1128/AEM.71.3.1553-1561.2005

M. J. Uriz, X. Turon, and M. A. Becerro, Morphology and ultrastructure of the swimming larvae of Crambe crambe (Demospongiae, Poecilosclerida), Invertebrate Biology, vol.226, issue.4, pp.295-307, 2001.
DOI : 10.1007/978-3-642-70892-3_15

S. Yooseph, Genomic and functional adaptation in surface ocean planktonic prokaryotes, Nature, vol.34, issue.7320, pp.60-66, 2010.
DOI : 10.1016/S0022-2836(05)80360-2

URL : http://www.nature.com/nature/journal/v468/n7320/pdf/nature09530.pdf

A. S. Peek, R. C. Vrijenhoek, and B. S. Gaut, Accelerated Evolutionary Rate in Sulfur-Oxidizing Endosymbiotic Bacteria Associated with the Mode of Symbiont Transmission, Molecular Biology and Evolution, vol.15, issue.11, pp.1514-1523, 1998.
DOI : 10.1093/oxfordjournals.molbev.a025879

N. A. Moran, Tracing the evolution of gene loss in obligate bacterial symbionts, Current Opinion in Microbiology, vol.6, issue.5, pp.512-518, 2003.
DOI : 10.1016/j.mib.2003.08.001

M. T. Suzuki, O. Beja, L. T. Taylor, and E. Delong, Phylogenetic analysis of ribosomal RNA operons from uncultivated coastal marine bacterioplankton, Environmental Microbiology, vol.63, issue.5, pp.323-331, 2001.
DOI : 10.1128/AEM.66.11.4605-4614.2000

N. Bensaadi-merchermek, J. C. Salvado, C. Cagnon, S. Karama, and C. Mouchès, Characterization of the unlinked 16S rDNA and 23S-5S rRNA operon of Wolbachia pipientis, a prokaryotic parasite of insect gonads, Gene, vol.165, issue.1, pp.81-86, 1995.
DOI : 10.1016/0378-1119(95)00539-I

F. R. Rurangirwa, K. A. Brayton, T. C. Mcguire, D. P. Knowles, and G. H. Palmer, Conservation of the unique rickettsial rRNA gene arrangement in Anaplasma, IJSEM, vol.52, pp.1405-1409, 2002.

K. M. Usher, J. Kuo, J. Fromont, and D. C. Sutton, Vertical transmission of cyanobacterial symbionts in the marine sponge Chondrilla australiensis (Demospongiae), Hydrobiologia, vol.461, issue.1/3, pp.9-13, 2001.
DOI : 10.1023/A:1012792426770

K. H. Sharp, B. Eam, D. J. Faulkner, and M. G. Haygood, Vertical Transmission of Diverse Microbes in the Tropical Sponge Corticium sp., Applied and Environmental Microbiology, vol.73, issue.2, pp.622-629, 2007.
DOI : 10.1128/AEM.01493-06

S. Schmitt, H. Angermeier, R. Schiller, N. Lindquist, and U. Hentschel, Molecular Microbial Diversity Survey of Sponge Reproductive Stages and Mechanistic Insights into Vertical Transmission of Microbial Symbionts, Applied and Environmental Microbiology, vol.74, issue.24, pp.7694-7708, 2008.
DOI : 10.1128/AEM.00878-08

J. Selvin, A. S. Ninawe, G. Seghal-kiran, and A. P. Lipton, Sponge-microbial interactions: Ecological implications and bioprospecting avenues, Critical Reviews in Microbiology, vol.68, issue.1, pp.82-90, 2010.
DOI : 10.1128/MMBR.68.4.745-770.2004

H. Bretting and K. Königsmann, Investigations on the lectin-producing cells in the sponge Axinella polypoides (Schmidt), Cell and Tissue Research, vol.201, issue.3, pp.487-497, 1979.
DOI : 10.1007/BF00237007

J. E. Thompson, K. D. Barrow, and D. J. Faulkner, ), Acta Zoologica, vol.16, issue.Suppl. 28, pp.199-210, 1983.
DOI : 10.1007/978-3-0348-5797-0

L. Fan, Functional equivalence and evolutionary convergence in complex communities of microbial sponge symbionts, Proceedings of the National Academy of Sciences, vol.587, issue.17, pp.1878-1887, 2012.
DOI : 10.1113/jphysiol.2009.174136

A. Siegl, Single-cell genomics reveals the lifestyle of Poribacteria, a candidate phylum symbiotically associated with marine sponges, The ISME Journal, vol.1784, issue.1, pp.61-70, 2011.
DOI : 10.1038/nbt1214

N. S. Webster, K. J. Wilson, L. L. Blackall, and R. T. Hill, Phylogenetic Diversity of Bacteria Associated with the Marine Sponge Rhopaloeides odorabile, Applied and Environmental Microbiology, vol.67, issue.1, pp.434-444, 2001.
DOI : 10.1128/AEM.67.1.434-444.2001

K. L. Vergin, Screening of a Fosmid Library of Marine Environmental Genomic DNA Fragments Reveals Four Clones Related to Members of the OrderPlanctomycetales, Appl Environ Microbiol, vol.64, pp.3075-3078, 1998.

G. Muyzer, Denaturing gradient gel electrophoresis (DGGE) in microbial ecology, pp.1-27, 1998.

J. K. Bonfield, K. F. Smith, and R. Staden, A new DNA sequence assembly program, Nucleic Acids Research, vol.23, issue.24, pp.4992-4999, 1995.
DOI : 10.1093/nar/23.24.4992

URL : https://academic.oup.com/nar/article-pdf/23/24/4992/7123392/23-24-4992.pdf

R. C. Edgar, B. J. Haas, J. C. Clemente, C. Quince, and R. Knight, UCHIME improves sensitivity and speed of chimera detection, Bioinformatics, vol.27, issue.16, pp.2194-2200, 2011.
DOI : 10.1093/bioinformatics/btr381

T. Huber, G. Faulkner, and P. Hugenholtz, Bellerophon: a program to detect chimeric sequences in multiple sequence alignments, Bioinformatics, vol.20, issue.14, pp.2317-2319, 2004.
DOI : 10.1093/bioinformatics/bth226

P. D. Schloss, Introducing mothur: Open-Source, Platform-Independent, Community-Supported Software for Describing and Comparing Microbial Communities, Applied and Environmental Microbiology, vol.75, issue.23, pp.7537-7541, 2009.
DOI : 10.1128/AEM.01541-09

URL : http://aem.asm.org/content/75/23/7537.full.pdf

C. Quast, The SILVA ribosomal RNA gene database project: improved data processing and web-based tools, Nucleic Acids Research, vol.41, issue.D1, pp.590-596, 2013.
DOI : 10.1093/nar/gks1219

URL : https://academic.oup.com/nar/article-pdf/41/D1/D590/3690367/gks1219.pdf

W. Ludwig, ARB: a software environment for sequence data, Nucleic Acids Research, vol.32, issue.4, pp.1363-1371, 2004.
DOI : 10.1093/nar/gkh293

J. Felsenstein and . Phylip, Phylogeny Inference Package) version 3.6. Distributed by the author, 2005.

F. Ronquist, MrBayes 3.2: Efficient Bayesian Phylogenetic Inference and Model Choice Across a Large Model Space, Systematic Biology, vol.61, issue.3, pp.539-542, 2012.
DOI : 10.1093/sysbio/sys029

S. E. Dowd, Polymicrobial Nature of Chronic Diabetic Foot Ulcer Biofilm Infections Determined Using Bacterial Tag Encoded FLX Amplicon Pyrosequencing (bTEFAP), PLoS ONE, vol.188, issue.3, p.3326, 2008.
DOI : 10.1371/journal.pone.0003326.t001

C. Quince, A. Lanzen, R. J. Davenport, and P. J. Turnbaugh, Removing Noise From Pyrosequenced Amplicons, BMC Bioinformatics, vol.12, issue.1, p.38, 2011.
DOI : 10.1186/1471-2105-12-38

URL : https://bmcbioinformatics.biomedcentral.com/track/pdf/10.1186/1471-2105-12-38?site=bmcbioinformatics.biomedcentral.com

J. G. Caporaso, QIIME allows analysis of high-throughput community sequencing data, Nature Methods, vol.8, issue.5, pp.335-336, 2010.
DOI : 10.1038/nmeth.f.303

R. C. Edgar, Search and clustering orders of magnitude faster than BLAST, Bioinformatics, vol.26, issue.19, pp.2460-2461, 2010.
DOI : 10.1093/bioinformatics/btq461

URL : https://academic.oup.com/bioinformatics/article-pdf/26/19/2460/16896486/btq461.pdf

B. M. Fuchs, F. O. Glöckner, J. Wulf, and R. Amann, Unlabeled Helper Oligonucleotides Increase the In Situ Accessibility to 16S rRNA of Fluorescently Labeled Oligonucleotide Probes, Applied and Environmental Microbiology, vol.66, issue.8, pp.3603-3607, 2000.
DOI : 10.1128/AEM.66.8.3603-3607.2000

S. Sauzet, Cloning and retinal expression of melatonin receptors in the European sea bass, Dicentrarchus labrax, General and Comparative Endocrinology, vol.157, issue.2, pp.186-195, 2008.
DOI : 10.1016/j.ygcen.2008.04.008

A. Pernthaler, J. Pernthaler, and R. Amann, Fluorescence In Situ Hybridization and Catalyzed Reporter Deposition for the Identification of Marine Bacteria, Applied and Environmental Microbiology, vol.68, issue.6, pp.3094-3101, 2002.
DOI : 10.1128/AEM.68.6.3094-3101.2002

M. J. Uriz, X. Turon, and M. A. Becerro, Silica deposition in Demosponges: spiculogenesis in Crambe crambe, Cell and Tissue Research, vol.301, issue.2, pp.299-309, 2000.
DOI : 10.1007/s004410000234

R. Amann, Combination of 16S rRNA-targeted oligonucleotide probes with flow cytometry for analyzing mixed microbial populations, Appl Environ Microbiol, vol.56, pp.1919-1925, 1990.

G. Wallner, R. Amann, and W. Beisker, Optimizing fluorescent in situ hybridization with rRNA-targeted oligonucleotide probes for flow cytometric identification of microorganisms, Cytometry, vol.137, issue.2, pp.136-143, 1993.
DOI : 10.1099/00221287-137-12-2823

W. Manz, R. Amann, W. Ludwig, and K. H. Schleifer, Phylogenetic Oligodeoxynucleotide Probes for the Major Subclasses of Proteobacteria: Problems and Solutions, Systematic and Applied Microbiology, vol.15, issue.4, pp.593-600, 1992.
DOI : 10.1016/S0723-2020(11)80121-9

D. A. Stahl and R. Amann, Development and application of nucleic acid probes, Nucleic acid techniques in bacterial systematics, pp.205-248, 1991.