L. R. Pastorino, D. Simone, and . Cavallari, ) helped complete our dataset We are grateful to Richard Preece (University Museum of Zoology, Cambridge) and Fred Naggs (The Natural History Museum, London) for sharing photos of the syntype of Clea annesleyi; to Manuel Caballer (E-RECOLNAT ANR-11-INBS-0004, MNHN) for photographing the types; to Delphine Gey (MNHN) for her efforts in optimizing sequencing protocols for freshwater Nassariidae; to Nicolas Puillandre (MNHN) for informative discussions and on the molecular aspects of the study; and to Philippe Bouchet (MNHN) who made this study possible and provided thoughtful advice and constructive feedback, especially on aspects related to nomenclature and classification. ADDITIONAL INFORMATION AND DECLARATIONS Funding This project was supported by the network " Bibliothèque du Vivant " funded by the CNRS-MNHN-INRA-CEA (Genoscope), the " Service de Systématique Moléculaire LAG received a PhD studentship from FUNDAYACUCHO-Venezuela and Campus France. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript, UMS 2700 CNRS-MNHN), the deep-sea studies developed in the French?Taiwanese project TF-DeepEvo ANR-NSC (ANR 12-ISV7-0005-01) and the Russian Science Foundation Grant Disclosures The following grant information was disclosed by the authors: Bibliothèque du Vivant: CNRS-MNHN-INRA-CEA

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