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, We thank L. Blanchoin's Cytomorpholab in Grenoble for providing us RPE1 cells and H. Maiato from the University of Porto for providing us with NIH-3T3 cells. Furthermore, we want to thank A. Kyumurkov at the Institute of Advanced Biology in Grenoble for assisting with adhesion imaging, Acknowledgments: We thank M. Piel and P. Maiuri for valuable discussions and sharing the cell race data with us. We also thank T. Boudou and M. Théry for critical discussions, J. Bernard for technical assistance, and members of the MOTIV team at LiPhy for support

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